Cyperus difformis |
Cyperus drummondii |
|
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Asian flatsedge, smallflower umbrella sedge, variable flatsedge |
Drummond's sedge |
|
Habit | Herbs, annual, cespitose. | Herbs, perennial, cespitose; rhizomes 0.5–2 cm, often absent. |
Culms | 1–15, trigonous, 7–30 cm × 1.2–2.5 mm, soft (flattened in pressing), glabrous. |
trigonous, 35–170 cm × 2–4.7 mm, scabrid on angles. |
Leaves | 2–7, flat, (2–)7–22 cm × 2.2–4 mm. |
4–12, with readily visible cross ribs especially on abaxial surface, flat to V-shaped, 25–100 cm × 2.4–11 mm. |
Inflorescences | heads dense, 7–17 mm diam.; when rays short, heads sessile or nearly so, then densely irregularly lobate, 12–35 mm diam.; rays 1–5, 2–32 mm; bracts 2–4, longest bract erect or nearly so, appearing as continuation of culm, other bracts horizontal to ascending, 1–22 cm × 0.5–3.5 mm, margins and keel minutely scabridulous. |
heads digitate, globose, 1–1.7 cm diam.; rays 3–5, 1–7 cm; 2d order rays absent; bracts 3–5, horizontal to ascending at 30°, with prominent cross ribs, V-shaped, (4–)11–35(–48) cm × 1–6.7 mm (longest 1/2 as long as culm); 2d order bracts absent. |
Spikelets | 30–120, greenish brown to purplish brown, oblong-ellipsoid, compressed, (2–)3–5(–6) × 0.8–1.2 mm; floral scales (6–)12–20(–30), laterally clear margins, stramineous to deep purple, medially greenish, stramineous, or purplish, laterally ribless, medially 3-ribbed, obovate to orbiculate, 0.6–0.8 × 0.6–0.8 mm, apex mucronulate. |
10–40, oblong to linear-lanceoloid, 5–1.8 × 1.5–2.2 mm; floral scales 18–42, pale grayish green, turning brownish, proximally greenish or brownish, 2-keeled, weakly to distinctly 1-ribbed, proximal 1/2 2-ribbed, ovate, 1.4–1.6 × 0.9–1.4 mm, acute (sometimes mucronulate), apically glabrous, occasionally scabridulous. |
Flowers | stamens 1 or 2; anthers ovoid-ellipsoid, 0.1 mm, connective not prolonged; styles 0.1 mm; stigmas 0.1–0.3 mm. |
stamens 1–2; anthers 0.8–1.2 mm; styles 0.8–1 mm; stigmas 0.6–0.8 mm. |
Achenes | light brown, obovoid-ellipsoid, 0.6–0.8 × 0.3–0.4 mm (as long as subtending scale), base cuneate, apex obtuse, apiculate, surfaces finely reticulate, papillose. |
brown, slightly stipitate, oblong-ellipsoid (2.4–3.9 times longer than wide), 1–1.2 × 0.3–0.5 mm, apical beak 0.1–0.5 mm, surfaces glabrous. |
Cyperus difformis |
Cyperus drummondii |
|
Phenology | Fruiting summer. | Fruiting late spring–summer. |
Habitat | Disturbed, muddy soils, shallow waters | Wet, relatively undisturbed habitats, especially flatwoods, ponds, seepage slopes, coastal prairies |
Elevation | 0–1000 m (0–3300 ft) | 0–200 m (0–700 ft) |
Distribution |
AL; AZ; CA; FL; GA; ID; KY; LA; MS; NC; NJ; NM; OR; PA; SC; TN; TX; VA; Mexico; South America; West Indies (Puerto Rico); Central America (Nicaragua, Panama); Eurasia; Indian Ocean Islands; Pacific Islands; Australia [Introduced in North America]
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AL; FL; GA; LA; MS; SC; TX; Central America; South America; West Indies |
Discussion | Cyperus difformis is naturalized in the New World and native to the Old World, where it ranges from southern Europe to southern Africa and eastward to Southeast Asia and Australia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus drummondii has been treated as a synonym (G. C. Tucker 1994), or a variety (G. Kükenthal 1935–1936), or a subspecies of C. virens (M. F. Denton 1978b). Recent quantitative and field studies (J. R. Carter et al. 1999) provided compelling evidence that specific status is appropriate. Compared to Cyperus virens, C. drummondii is a taller species of less disturbed habitats and has narrower spikelets, ovate scales, and fewer inflorescence bracts. Cyperus virens and C. drummondii are readily distinguished from other Cyperus species in the United States by their sharply angled, scabrid culms and conspicuously septate leaf blades. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 156. | FNA vol. 23. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Pycnostachys |
Sibling taxa | ||
Synonyms | C. lateriflorus | C. robustus, C. virens subsp. drummondii, C. virens var. drummondii, C. virens var. robustus |
Name authority | Linnaeus: Cent. Pl. II, 6. (1756) | Torrey & Hooker: Ann. Lyceum Nat. Hist. New York 3: 437, errata. (1836) |
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