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Asian flatsedge, smallflower umbrella sedge, variable flatsedge

cyperus, flatsedge, galingale, nutsedge, umbrella-sedge

Habit Herbs, annual, cespitose. Herbs, perennial or less often annual, cespitose or not, rhizomatous, stoloniferous, rarely tuberous.
Culms

1–15, trigonous, 7–30 cm × 1.2–2.5 mm, soft (flattened in pressing), glabrous.

solitary or not, trigonous or round, glabrous or scabridulous with extrorse or antrorse (rarely retrorse) prickles.

Leaves

2–7, flat, (2–)7–22 cm × 2.2–4 mm.

usually basal;

ligules absent;

blades keeled abaxially, flat, V-, or inversely W-shaped in cross section.

Inflorescences

heads dense, 7–17 mm diam.; when rays short, heads sessile or nearly so, then densely irregularly lobate, 12–35 mm diam.;

rays 1–5, 2–32 mm;

bracts 2–4, longest bract erect or nearly so, appearing as continuation of culm, other bracts horizontal to ascending, 1–22 cm × 0.5–3.5 mm, margins and keel minutely scabridulous.

terminal, rarely pseudolateral, 1st order subumbellate to capitate, 2d order with spicate or digitately arranged spikelets, rarely a solitary spikelet;

spikelets 1–150; 1st order rays unequal (rarely equal) in length, produced singly from the axils of inflorescence bracts;

involucral bracts 1–22, spirally arranged at culm apex, spreading to erect, leaflike.

Spikelets

30–120, greenish brown to purplish brown, oblong-ellipsoid, compressed, (2–)3–5(–6) × 0.8–1.2 mm;

floral scales (6–)12–20(–30), laterally clear margins, stramineous to deep purple, medially greenish, stramineous, or purplish, laterally ribless, medially 3-ribbed, obovate to orbiculate, 0.6–0.8 × 0.6–0.8 mm, apex mucronulate.

scales to 76, distichous, each subtending flower, cylindric to compressed, borne spicately or digitately at ends of rays (occasionally proliferous).

Flowers

stamens 1 or 2;

anthers ovoid-ellipsoid, 0.1 mm, connective not prolonged;

styles 0.1 mm;

stigmas 0.1–0.3 mm.

bisexual [rarely unisexual], in axils of distichous floral scales, bases often decurrent onto rachilla as ± hyaline wings;

perianth absent;

stamens 1–3;

styles linear, 2–3-fid, base deciduous or persistent;

stigmas 2–3.

Achenes

light brown, obovoid-ellipsoid, 0.6–0.8 × 0.3–0.4 mm (as long as subtending scale), base cuneate, apex obtuse, apiculate, surfaces finely reticulate, papillose.

biconvex, flattened, or trigonous.

Cyperus difformis

Cyperus

Phenology Fruiting summer.
Habitat Disturbed, muddy soils, shallow waters
Elevation 0–1000 m (0–3300 ft)
Distribution
from FNA
AL; AZ; CA; FL; GA; ID; KY; LA; MS; NC; NJ; NM; OR; PA; SC; TN; TX; VA; Mexico; South America; West Indies (Puerto Rico); Central America (Nicaragua, Panama); Eurasia; Indian Ocean Islands; Pacific Islands; Australia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from USDA
Pantemperate and tropical
[BONAP county map]
Discussion

Cyperus difformis is naturalized in the New World and native to the Old World, where it ranges from southern Europe to southern Africa and eastward to Southeast Asia and Australia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species ca. 600 (96 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stigmas 2 (3 in C. serotinus); achenes biconvex.
→ 2
1. Stigmas 3; achenes trigonous, plano-convex, or terete.
→ 3
2. Achenes laterally flattened, borne with edge toward rachilla.
subg. Pycreus
2. Achenes dorsiventrally flattened, borne with face toward rachilla.
subg. Juncellus
3. Spikelets borne in digitate clusters (rarely singly) or in umbellate or glomerulate heads.
subg. Pycnostachys
3. Spikelets borne in spikes on conspicuous rachis.
→ 4
4. Rachilla continuous or articulate only at base.
subg. Cyperus
4. Rachilla articulate at base of each scale, mature spikelet disarticulating into segments consisting of scale, internode, and rachilla wings.
subg. Diclidium
Source FNA vol. 23, p. 156. FNA vol. 23, p. 141. Authors: Gordon C. Tucker*, Brian G. Marcks*, J. Richard Carter *.
Parent taxa Cyperaceae > Cyperus > subg. Pycnostachys Cyperaceae
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Subordinate taxa
C. subg. Cyperus, C. subg. Diclidium, C. subg. Juncellus, C. subg. Pycnostachys, C. subg. Pycreus
Synonyms C. lateriflorus
Name authority Linnaeus: Cent. Pl. II, 6. (1756) Linnaeus: Sp. Pl. 1: 44. (1753): Gen. Pl. ed. 5, 26. (1754)
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