Cylindropuntia whipplei |
Cylindropuntia munzii |
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Whipple cholla |
Munz cholla, Munz's cholla |
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Habit | Trees or shrubs, low to upright, sparingly to densely branched, some almost matlike in Arizona, 0.3–0.6(–1.5) m; branchlets whorled to subwhorled. | Trees or treelike shrubs, with trunk(s) and main branches spreading, bearing terminal tufts of usually drooping branchlets, 2–4 m. Stem segments easily detached, gray-green, 4–16 × 1.2–2.5 cm; tubercles prominent, narrowly oval, 1–2 cm; areoles subcircular to obdeltate, 5–7 × 3–4.5 mm; wool tawny to yellow-brown, aging gray. |
Stem | segments firmly attached, green, 3–9(–15) × 0.5–1.5(–2.2) cm; tubercles prominent, short, 0.5–1 cm; areoles oval to obdeltate, 2–6 × 1.5–4 mm; wool pale yellow to white, aging gray. |
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Spines | (1–)3–8(–10) spines per areole, in all but basalmost areoles, best developed toward apex, interlacing with spines of adjacent areoles, whitish or pale yellow, pale red-brown, sometimes tipped yellow, of 2 kinds; radial spines slender, flattened basally, deflexed, 5–8 mm, surrounding central spines; central spines usually 4(–6), spreading into a cross, stout, subterete, 2–3.4(–4.5) cm; also 0–2 bristlelike spines; sheaths whitish to pale yellow (rarely golden) throughout or tipped yellow to golden. |
(7–)9–14 per areole, at most areoles, erect or spreading, yellow, aging red- to gray-black with pale tips, terete or abaxial ones slightly flattened, the longest to 3 cm; marginal bristlelike spines shorter, 0–4; sheaths pale yellow becoming golden apically, baggy. |
Glochids | in adaxial tuft, yellow, 1–3 mm. |
in adaxial crescent and sometimes small tuft, yellow, inconspicuous, 0.5–2 mm. |
Flowers | inner tepals yellow to green-yellow, spatulate, 15–25(–30) mm, apiculate; filaments yellowish or yellow-green; anthers yellow; style white to yellowish; stigma lobes whitish, yellowish, or pale green (rarely pink tinged). |
inner tepals pale reddish maroon-brown, elliptic, of irregular lengths, 8–15 mm, emarginate-apiculate; filaments green; anthers yellow; style and stigma lobes cream. |
Fruits | rarely proliferating, yellow to greenish yellow, broadly cylindric to subspheric, 18–30(–35) × 15–22(–32) mm, fleshy, tuberculate, spineless; tubercles subequal or distalmost longest, usually prominent; umbilicus 7–8 mm deep; areoles 36–62. |
easily detached, tan when mature and fertile, globose, 17–24 × 17–21 mm, dry, tuberculate, spineless but bearing numerous very long glochids; basal tubercles not markedly longer than distal ones (except in sterile fruits); areoles 30–40. |
Seeds | pale yellow, subcircular to slightly angular in outline, flattened to warped, 3–3.5 × 2.5–3.5 mm, sides smooth or with 1–3 depressions; girdle smooth. |
pale yellow, rounded-deltoid, slightly flattened, 3–4.5 × 3–4 mm, sides smooth; girdle smooth, not protruding. |
2n | = 22, 44. |
= 22, 33. |
Cylindropuntia whipplei |
Cylindropuntia munzii |
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Phenology | Flowering late spring–early summer (May–Jul). | Flowering spring (Mar–May). |
Habitat | Desert and plains grasslands, juniper woodlands, oak, pinyon, or pine forests, sagebrush | Sonoran Desert, flats, hills, sandy to rocky soils |
Elevation | 900-2300 m (3000-7500 ft) | 400-700 m (1300-2300 ft) |
Distribution |
AZ; CO; NM; NV; UT
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CA; Mexico (Baja California) |
Discussion | Cylindropuntia whipplei hybridizes with C. acanthocarpa in central Arizona; the hybrids [= C. ×congesta (Griffiths) F. M. Knuth (as species) (D. J. Pinkava 1999)] are compact shrubs with long-tuberculate stem segments 8–12 mm thick, bearing 0–4 erect or deflexed spines per areole, and tuberculate, green to yellowish fruits occasionally bearing one to few spines, and have chromosome number of 2n = 22. In areas of Arizona, Nevada, and Utah, C. whipplei hybridizes with C. echinocarpa, particularly with the low, compact form of C. echinocarpa; the hybrids [= C. ×multigeniculata (Clokey) Backeberg] have spineless to nearly spineless fruits, numerous intergrading spines per stem areole that are not dimorphic, and chromosome number of 2n = 22. Hybrids between C. whipplei and C. imbricata [= C. ×viridiflora (Britton & Rose) F. M. Knuth], which are found in scattered localities in northern New Mexico and northeastern Arizona, have a low, bushy habit, ripe cantaloupe-colored inner tepals, an irregular, monomorphic spine pattern, and a chromosome number of 2n = 22. Cylindropuntia ×media P. V. Heath, from the base of Wolf Creek Pass, Colorado (G. Arp 1973), reportedly has the same putative parentage as the older name C. ×viridiflora; further study of its parentage is warranted. Cylindropuntia whipplei also hybridizes with C. leptocaulis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cylindropuntia munzii hybridizes with C. echinocarpa forming a shrub to 1.5 m, with shorter tubercles and baggy-sheathed spines obscuring the stem and bearing fruits with long glochids but no spines. The chromosome number for the hybrid is 2n = 22. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 4, p. 111. | FNA vol. 4, p. 115. |
Parent taxa | Cactaceae > subfam. Opuntioideae > Cylindropuntia | Cactaceae > subfam. Opuntioideae > Cylindropuntia |
Sibling taxa | ||
Synonyms | Opuntia whipplei, C. hualpaensis, C. whipplei var. enodis, Opuntia whipplei var. enodis, Opuntia whipplei var. laevior | Opuntia munzii |
Name authority | (Engelmann & J. M. Bigelow) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 124. (1935) | (C. B. Wolf) Backeberg: Kakteenlexikon, 113. (1966) |
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