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staghorn cholla

kelvin pricklypear

Habit Trees or shrubs, openly branching at acute angles, to 2 m. Stem segments whorled or subwhorled, purple to green-purple, 4–18 × 1–2 cm; tubercles prominent, elongate-oval, 1–2(–2.5) cm; areoles subcircular, 3.5–4 mm diam.; wool tan to brown, aging gray. Trees 1.5–2 m; trunk commonly branched; crown open to compact usually bearing whorled branches.
Stem

segments often easily dislodged, if terminal, green or purple, 5–13 × 1.8–3 cm;

tubercles prominent, broadly oval, 0.7–1.2(–1.5) cm;

areoles obdeltate to rhombic, 4.5–6 × 3–5 mm;

wool yellow, aging gray to black.

Spines

6–8 per areole (1–2 bristlelike spines) increasing with time, well distributed along stem, slightly interlacing with spines of adjacent areoles, short;

abaxial spines usually reflexed, whitish, pinkish to red-brown, lightly flattened, the longest 10–18 mm;

adaxial spines erect or spreading, rich red-brown, gray coated basally, subterete, 6–11 mm;

sheaths grayish, tipped brown or yellow to golden Glochids in small adaxial tuft or crescent, dark yellow, to 1 mm.

0–8(–13) per areole, sometimes with 1–3 bristlelike spines along areole margins at most areoles, usually slightly interlacing with spines of adjacent areoles, yellowish to tan, sometimes with pinkish hue, to red-brown and partially gray coated, aging brown-gray to gray;

major abaxial spines angular-flattened, usually reflexed, the longest to 30 mm;

major adaxial spines subterete, erect-divergent, the longest to 18 mm;

sheaths uniformly whitish or tipped yellow to gold, slightly baggy.

Glochids

in inconspicuous adaxial crescent, yellow, to 1 mm.

Flowers

inner tepals yellow-green, yellow to gold or bronze, or red to rose or magenta, spatulate, apiculate;

filaments yellowish green;

anthers yellow;

style whitish to pale bronze;

stigma lobes whitish.

inner tepals rose to magenta, spatulate, 18–25(–30) mm, emarginate-apiculate;

filaments deep rose;

anthers yellow;

style white, tinged red-purple distally;

stigma lobes white with hint of green.

Fruits

often proliferating, yellowish green or tinged red to purple, often stipitate, obovate, 25–40 × 10–20 mm, leathery-fleshy, tuberculate or smooth, essentially spineless;

tubercles prominent, at first, subequal in length or proximal ones longer, fertile ones swelling and becoming smooth;

umbilicus to 9 mm deep;

areoles 20–30.

usually sterile, sometimes forming short chains, yellow-green to yellow, sometimes purplish, 25–45 × 15–22 mm, fleshy, tuberculate, spineless;

tubercles progressively longer toward fruit apex;

umbilicus 4–7 mm deep;

areoles 32–44.

Seeds

yellowish, polygonal in outline, warped, 3.5–5.5 × 2.5–5 mm, sides nearly flattened with depressions and protrusions to angular subspheric;

girdle smooth or in a groove.

pale yellow, orbicular to angular or squarish in outline, flattened to warped, 3.5–4 × 3–4 mm, sides with 0–2 large depressions;

hilum sometimes pointed;

girdle smooth.

2n

= 22.

= 22, 33.

Cylindropuntia versicolor

Cylindropuntia ×kelvinensis

Phenology Flowering spring (Apr–Jun). Flowering spring (Apr–Jun).
Habitat Sonoran Desert, desert scrub, flats, washes, rocky hillsides, canyons Sonoran desert scrub, edges of grasslands, rocky flats and slopes, rolling hills
Elevation 600-1300 m (2000-4300 ft) 500-1000 m (1600-3300 ft)
Distribution
from FNA
AZ; Mexico (Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ
[BONAP county map]
Discussion

Cylindropuntia versicolor forms hybrids with C. acanthocarpa var. major, C. arbuscula (= C. ×vivipara), C. leptocaulis (see discussion under 3. C. ×tetracantha), and C. spinosior (= C. ×grantiorum P. V. Heath). Cylindropuntia ×grantiorum, found in eastern to south-central Arizona, is intermediate between the parental species in values for the number of spines per stem areole, stem segment diameter and color, and fruit tubercles. The hybrids between C. versicolor and C. acanthocarpa var. major, which occur in south-central Arizona, have fleshy, tuberculate fruits, often with one or two short permanent spines at apex and longer and more numerous spines per stem areole.

Cylindropuntia versicolor itself may be of hybrid origin.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 4, p. 109. FNA vol. 4, p. 108.
Parent taxa Cactaceae > subfam. Opuntioideae > Cylindropuntia Cactaceae > subfam. Opuntioideae > Cylindropuntia
Sibling taxa
C. abyssi, C. acanthocarpa, C. arbuscula, C. bigelovii, C. californica, C. davisii, C. echinocarpa, C. fulgida, C. ganderi, C. imbricata, C. kleiniae, C. leptocaulis, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. whipplei, C. wolfii, C. ×kelvinensis, C. ×tetracantha
C. abyssi, C. acanthocarpa, C. arbuscula, C. bigelovii, C. californica, C. davisii, C. echinocarpa, C. fulgida, C. ganderi, C. imbricata, C. kleiniae, C. leptocaulis, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. versicolor, C. whipplei, C. wolfii, C. ×tetracantha
Synonyms Opuntia versicolor, Opuntia arborescens var. versicolor, Opuntia thurberi subsp. versicolor Opuntia kelvinensis
Name authority (Engelmann ex J. M. Coulter) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 125. (1935) (V. E. Grant & K. A. Grant) P. V. Heath: Calyx 4: 142. (1994)
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