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staghorn cholla

candle cholla, Klein cholla, Klein's pencil cactus, tasajillo

Habit Trees or shrubs, openly branching at acute angles, to 2 m. Stem segments whorled or subwhorled, purple to green-purple, 4–18 × 1–2 cm; tubercles prominent, elongate-oval, 1–2(–2.5) cm; areoles subcircular, 3.5–4 mm diam.; wool tan to brown, aging gray. Shrubs scraggly, openly branched, 0.5–2.5 m. Stem segments usually alternate, green, 4–20 × 0.6–1.2 cm; tubercles obvious, 1–2.5 cm; areoles subcircular to oval, 2.5–3.5 × 2–3 mm; wool yellow, aging gray to black.
Spines

6–8 per areole (1–2 bristlelike spines) increasing with time, well distributed along stem, slightly interlacing with spines of adjacent areoles, short;

abaxial spines usually reflexed, whitish, pinkish to red-brown, lightly flattened, the longest 10–18 mm;

adaxial spines erect or spreading, rich red-brown, gray coated basally, subterete, 6–11 mm;

sheaths grayish, tipped brown or yellow to golden Glochids in small adaxial tuft or crescent, dark yellow, to 1 mm.

(0–)1–2(–4) per areole, at most areoles, straight to slightly arched, commonly deflexed, not obscuring stem, yellow to gray, tipped yellow, acicular, the longest 1–3 cm.

Glochids

in adaxial tuft, yellow, 0.5–2.5 mm.

Flowers

inner tepals yellow-green, yellow to gold or bronze, or red to rose or magenta, spatulate, apiculate;

filaments yellowish green;

anthers yellow;

style whitish to pale bronze;

stigma lobes whitish.

inner tepals greenish basally to reddish bronze, tinged magenta apically, spatulate-apiculate, 15–25 mm;

filaments greenish basally to bronze distally;

anthers yellow;

style greenish bronze;

stigma lobes cream.

Fruits

often proliferating, yellowish green or tinged red to purple, often stipitate, obovate, 25–40 × 10–20 mm, leathery-fleshy, tuberculate or smooth, essentially spineless;

tubercles prominent, at first, subequal in length or proximal ones longer, fertile ones swelling and becoming smooth;

umbilicus to 9 mm deep;

areoles 20–30.

green becoming red, obovoid to cylindric, 13–34 × 10–20 mm, fleshy, low tuberculate to smooth, spineless;

tubercles subequal;

umbilicus 3–6 mm deep;

areoles 16–28.

Seeds

yellowish, polygonal in outline, warped, 3.5–5.5 × 2.5–5 mm, sides nearly flattened with depressions and protrusions to angular subspheric;

girdle smooth or in a groove.

tan, squarish to oval, 4–5 × 3.5–4 mm, warped, sides smooth to lumpy;

girdle broad, low-ridged.

2n

= 22.

= 44.

Cylindropuntia versicolor

Cylindropuntia kleiniae

Phenology Flowering spring (Apr–Jun). Flowering late spring–early summer (May–Jun).
Habitat Sonoran Desert, desert scrub, flats, washes, rocky hillsides, canyons Chihuahuan Desert, creosote bush-mesquite flats, limestone rocky slopes
Elevation 600-1300 m (2000-4300 ft) 800-1800 m (2600-5900 ft)
Distribution
from FNA
AZ; Mexico (Sonora)
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[BONAP county map]
from FNA
NM; TX; Mexico (Chihuahua, Coahuila, Durango, Nuevo León)
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[BONAP county map]
Discussion

Cylindropuntia versicolor forms hybrids with C. acanthocarpa var. major, C. arbuscula (= C. ×vivipara), C. leptocaulis (see discussion under 3. C. ×tetracantha), and C. spinosior (= C. ×grantiorum P. V. Heath). Cylindropuntia ×grantiorum, found in eastern to south-central Arizona, is intermediate between the parental species in values for the number of spines per stem areole, stem segment diameter and color, and fruit tubercles. The hybrids between C. versicolor and C. acanthocarpa var. major, which occur in south-central Arizona, have fleshy, tuberculate fruits, often with one or two short permanent spines at apex and longer and more numerous spines per stem areole.

Cylindropuntia versicolor itself may be of hybrid origin.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In western Texas and northern Mexico, Cylindropuntia kleiniae forms hybrids with C. leptocaulis (= C. ×antoniae P. V. Heath) that have varying degrees of morphologic intermediacy and chromosome number of 2n = 33. In northern Mexico, C. kleiniae hybridizes with C. imbricata var. imbricata. Cylindropuntia kleiniae itself may be of hybrid origin.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 4, p. 109. FNA vol. 4, p. 107.
Parent taxa Cactaceae > subfam. Opuntioideae > Cylindropuntia Cactaceae > subfam. Opuntioideae > Cylindropuntia
Sibling taxa
C. abyssi, C. acanthocarpa, C. arbuscula, C. bigelovii, C. californica, C. davisii, C. echinocarpa, C. fulgida, C. ganderi, C. imbricata, C. kleiniae, C. leptocaulis, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. whipplei, C. wolfii, C. ×kelvinensis, C. ×tetracantha
C. abyssi, C. acanthocarpa, C. arbuscula, C. bigelovii, C. californica, C. davisii, C. echinocarpa, C. fulgida, C. ganderi, C. imbricata, C. leptocaulis, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. versicolor, C. whipplei, C. wolfii, C. ×kelvinensis, C. ×tetracantha
Synonyms Opuntia versicolor, Opuntia arborescens var. versicolor, Opuntia thurberi subsp. versicolor Opuntia kleiniae, Opuntia wrightii
Name authority (Engelmann ex J. M. Coulter) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 125. (1935) (de Candolle) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kactus-ABC, 123. (1935)
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