Cylindropuntia versicolor |
Cylindropuntia ganderi |
|
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staghorn cholla |
Gander cholla, Gander's buckhorn cholla, Gander's cholla |
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Habit | Trees or shrubs, openly branching at acute angles, to 2 m. Stem segments whorled or subwhorled, purple to green-purple, 4–18 × 1–2 cm; tubercles prominent, elongate-oval, 1–2(–2.5) cm; areoles subcircular, 3.5–4 mm diam.; wool tan to brown, aging gray. | Shrubs, ascending, with strict branches, 0.5–1.5 m. Stem segments firmly attached, green, (5–)10–26 × (1.7–)2.5–4.5 cm; tubercles prominent, narrowly elliptic to oval, often subribbed, (0.9–)1.3–2.6 cm; areoles elliptic to obdeltate, 4–7 × 2.5–5 mm; wool yellowish tan, aging gray. |
Spines | 6–8 per areole (1–2 bristlelike spines) increasing with time, well distributed along stem, slightly interlacing with spines of adjacent areoles, short; abaxial spines usually reflexed, whitish, pinkish to red-brown, lightly flattened, the longest 10–18 mm; adaxial spines erect or spreading, rich red-brown, gray coated basally, subterete, 6–11 mm; sheaths grayish, tipped brown or yellow to golden Glochids in small adaxial tuft or crescent, dark yellow, to 1 mm. |
11–18(–28) per areole, at most areoles, moderately to wholly obscuring stems, spreading or erect, whitish, pale yellow to pinkish tan to reddish, aging brown or black, terete or abaxial spines basally flattened, grading in length, the longest porrect to reflexed, 2–2.5(–3.5) cm, with 0–6 bristlelike spines at areole margins; sheaths slightly baggy, whitish becoming yellow to golden apically. |
Glochids | in thin adaxial crescent, yellow or tan to rusty, 0.5–6 mm. |
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Flowers | inner tepals yellow-green, yellow to gold or bronze, or red to rose or magenta, spatulate, apiculate; filaments yellowish green; anthers yellow; style whitish to pale bronze; stigma lobes whitish. |
inner tepals greenish yellow, often with tips reddish abaxially, spatulate, 22–30 mm, apiculate to emarginate-apiculate; filaments yellow-green to green; anthers yellow to orange-yellow; style cream to yellow, sometimes basally greenish; stigma lobes greenish cream, cream or yellow. |
Fruits | often proliferating, yellowish green or tinged red to purple, often stipitate, obovate, 25–40 × 10–20 mm, leathery-fleshy, tuberculate or smooth, essentially spineless; tubercles prominent, at first, subequal in length or proximal ones longer, fertile ones swelling and becoming smooth; umbilicus to 9 mm deep; areoles 20–30. |
yellowish, drying tan, globose, tuberculate, budlike with spines similar to those of stem segments but slightly shorter, often reflexed; proximal tubercles slightly elongated if fruits fertile; umbilicus 4–5 mm deep; areoles 22–32(–40). |
Seeds | yellowish, polygonal in outline, warped, 3.5–5.5 × 2.5–5 mm, sides nearly flattened with depressions and protrusions to angular subspheric; girdle smooth or in a groove. |
pale yellow, angular-suborbicular, flattened to slightly warped, 4.5–6 × 4.5–5.7 mm, sides with 3–5 depressions; girdle smooth, not protruding. |
2n | = 22. |
= 22. |
Cylindropuntia versicolor |
Cylindropuntia ganderi |
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Phenology | Flowering spring (Apr–Jun). | Flowering spring (Mar–May). |
Habitat | Sonoran Desert, desert scrub, flats, washes, rocky hillsides, canyons | Desert alluvial flats, bajadas, sandy, gravelly to rocky substrates |
Elevation | 600-1300 m (2000-4300 ft) | 100-1000 m (300-3300 ft) |
Distribution |
AZ; Mexico (Sonora)
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CA; Mexico (Baja California)
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Discussion | Cylindropuntia versicolor forms hybrids with C. acanthocarpa var. major, C. arbuscula (= C. ×vivipara), C. leptocaulis (see discussion under 3. C. ×tetracantha), and C. spinosior (= C. ×grantiorum P. V. Heath). Cylindropuntia ×grantiorum, found in eastern to south-central Arizona, is intermediate between the parental species in values for the number of spines per stem areole, stem segment diameter and color, and fruit tubercles. The hybrids between C. versicolor and C. acanthocarpa var. major, which occur in south-central Arizona, have fleshy, tuberculate fruits, often with one or two short permanent spines at apex and longer and more numerous spines per stem areole. Cylindropuntia versicolor itself may be of hybrid origin. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In the north part of its range, Cylindropuntia ganderi intergrades with C. californica var. parkeri. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 4, p. 109. | FNA vol. 4, p. 115. |
Parent taxa | Cactaceae > subfam. Opuntioideae > Cylindropuntia | Cactaceae > subfam. Opuntioideae > Cylindropuntia |
Sibling taxa | ||
Synonyms | Opuntia versicolor, Opuntia arborescens var. versicolor, Opuntia thurberi subsp. versicolor | Opuntia acanthocarpa subsp. ganderi, Opuntia acanthocarpa var. ganderi, Opuntia ganderi |
Name authority | (Engelmann ex J. M. Coulter) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 125. (1935) | (C. B. Wolf) Rebman & Pinkava: J. Arizona-Nevada Acad. Sci. 33: 150. (2001) |
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