Cylindropuntia versicolor |
Cylindropuntia echinocarpa |
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staghorn cholla |
golden cholla, silver cholla, silver or golden cholla, Wiggins' cholla |
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Habit | Trees or shrubs, openly branching at acute angles, to 2 m. Stem segments whorled or subwhorled, purple to green-purple, 4–18 × 1–2 cm; tubercles prominent, elongate-oval, 1–2(–2.5) cm; areoles subcircular, 3.5–4 mm diam.; wool tan to brown, aging gray. | Trees or shrubs, spreading, densely branched, 0.5–2 m. Stem segments usually firmly attached, terminal segments sometimes easily detached, green to light gray-green, cylindric, 3–7.5(–12) × 1–2.5 cm; tubercles salient, short oval, 0.4–1.3(–1.5) cm, moderately broad; areoles broadly elliptic, 4–4.5 × 2–3.5 mm; wool whitish, yellow to tan, aging gray. |
Spines | 6–8 per areole (1–2 bristlelike spines) increasing with time, well distributed along stem, slightly interlacing with spines of adjacent areoles, short; abaxial spines usually reflexed, whitish, pinkish to red-brown, lightly flattened, the longest 10–18 mm; adaxial spines erect or spreading, rich red-brown, gray coated basally, subterete, 6–11 mm; sheaths grayish, tipped brown or yellow to golden Glochids in small adaxial tuft or crescent, dark yellow, to 1 mm. |
(6–)10–17(–22) per areole, accompanied by 0–5 short bristlelike spines at areole margins, at most areoles, most prominent in adaxial ones, much interlaced with those of adjacent areoles, obscuring stems, sometimes brushlike; major abaxial spines whitish, yellow to tan, usually subterete to flattened, spreading to deflexed, the longest 20–30(–45) mm; major adaxial spines erect or spreading, white, pale yellow, or tan to brown (rarely deep red-brown), terete, the longest 22–35(–50) mm; sheaths whitish with yellow to golden tips, baggy. |
Glochids | conspicuous in broad adaxial crescent, yellow, 3–4(–5) mm. |
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Flowers | inner tepals yellow-green, yellow to gold or bronze, or red to rose or magenta, spatulate, apiculate; filaments yellowish green; anthers yellow; style whitish to pale bronze; stigma lobes whitish. |
inner tepals light green to yellow-green, sometimes suffused with maroon or rose, spatulate, 20–23 mm, emarginate-apiculate; filaments greenish white or yellow, sometimes suffused with bronze or rose, but not red; anthers yellow; style and stigma lobes whitish, cream to light green. |
Fruits | often proliferating, yellowish green or tinged red to purple, often stipitate, obovate, 25–40 × 10–20 mm, leathery-fleshy, tuberculate or smooth, essentially spineless; tubercles prominent, at first, subequal in length or proximal ones longer, fertile ones swelling and becoming smooth; umbilicus to 9 mm deep; areoles 20–30. |
tan at maturity, obconic to subspheric, 13–23 × 11–20 mm, dry, tuberculate, densely spiny, sometimes burlike, with apical flange around deep umbilicus; basal tubercles not markedly longer than distal ones; areoles 16–24(–36). |
Seeds | yellowish, polygonal in outline, warped, 3.5–5.5 × 2.5–5 mm, sides nearly flattened with depressions and protrusions to angular subspheric; girdle smooth or in a groove. |
pale yellow, angular to squarish in outline, warped, 4–6 × 3.5–5 mm, sides smooth, each with 2–4 large depressions; girdle smooth (rarely with very narrow ridge). |
2n | = 22. |
= 22. |
Cylindropuntia versicolor |
Cylindropuntia echinocarpa |
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Phenology | Flowering spring (Apr–Jun). | Flowering spring (Mar–Jun). |
Habitat | Sonoran Desert, desert scrub, flats, washes, rocky hillsides, canyons | Mojave and Sonoran deserts, desert grasslands, juniper and oak-juniper woodlands, flats, bajadas, canyons, sandy, loam, alluvial to gravelly substrates |
Elevation | 600-1300 m (2000-4300 ft) | 50-1700 m (200-5600 ft) |
Distribution |
AZ; Mexico (Sonora)
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AZ; CA; NV; Mexico (Baja California, Sonora)
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Discussion | Cylindropuntia versicolor forms hybrids with C. acanthocarpa var. major, C. arbuscula (= C. ×vivipara), C. leptocaulis (see discussion under 3. C. ×tetracantha), and C. spinosior (= C. ×grantiorum P. V. Heath). Cylindropuntia ×grantiorum, found in eastern to south-central Arizona, is intermediate between the parental species in values for the number of spines per stem areole, stem segment diameter and color, and fruit tubercles. The hybrids between C. versicolor and C. acanthocarpa var. major, which occur in south-central Arizona, have fleshy, tuberculate fruits, often with one or two short permanent spines at apex and longer and more numerous spines per stem areole. Cylindropuntia versicolor itself may be of hybrid origin. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
A dwarf form, with narrow terminal stem segments bearing few spines per areole, and one spine longer than the others, occurs on the very arid flats along the lower Colorado River in California and Arizona and has been recognized as Opuntia wigginsii L. D. Benson; however, numerous intermediate to more robust forms make recognition of the dwarf form untenable. Although exceptions occur, plants to the north and west in the range tend to be more spiny and to bear yellow to yellow-green flowers. Cylindropuntia echinocarpa hybridizes with C. acanthocarpa [= C. ×deserta (Griffiths) Pinkava (D. J. Pinkava 1999)], C. whipplei, and C. munzii. Cylindropuntia ×deserta, found in western Arizona and southern California, has reddish filaments, short tubercles, variably sized spines/areole, and dry spiny fruits. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 4, p. 109. | FNA vol. 4, p. 115. |
Parent taxa | Cactaceae > subfam. Opuntioideae > Cylindropuntia | Cactaceae > subfam. Opuntioideae > Cylindropuntia |
Sibling taxa | ||
Synonyms | Opuntia versicolor, Opuntia arborescens var. versicolor, Opuntia thurberi subsp. versicolor | Opuntia echinocarpa, Cactus echinocarpus, C. wigginsii, Opuntia wigginsii |
Name authority | (Engelmann ex J. M. Coulter) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 125. (1935) | (Engelmann & J. M. Bigelow) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 124. (1935) |
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