Cylindropuntia spinosior |
Cylindropuntia kleiniae |
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cane cholla, walking stick cholla, walkingstick cactus |
candle cholla, Klein cholla, Klein's pencil cactus, tasajillo |
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Habit | Trees, compact, widely branching, 0.4–2 m. Stem segments firmly attached, whorled or subwhorled, green to purple, 5–23 × 1.3–3.5 cm; tubercles crowded, pronounced, oval to narrowly oval, 0.5–1.2(–1.5) cm; areoles broadly obdeltate to elliptic, 4.5–7 × 2–4 mm; wool yellow to tan, aging gray to black. | Shrubs scraggly, openly branched, 0.5–2.5 m. Stem segments usually alternate, green, 4–20 × 0.6–1.2 cm; tubercles obvious, 1–2.5 cm; areoles subcircular to oval, 2.5–3.5 × 2–3 mm; wool yellow, aging gray to black. |
Spines | (4–)6–18(–24) per areole, at most areoles, interlacing with spines of adjacent areoles, pale tan (rarely yellowish), pinkish to red-brown; abaxial spines erect to usually deflexed, terete, often flattened basally, 8–19 mm; adaxial spines erect or spreading, subterete; ± bristlelike spines at areole abaxial margins; sheaths persisting, uniformly whitish, not baggy. |
(0–)1–2(–4) per areole, at most areoles, straight to slightly arched, commonly deflexed, not obscuring stem, yellow to gray, tipped yellow, acicular, the longest 1–3 cm. |
Glochids | in inconspicuous adaxial tuft, yellow to tan, aging gray, 1–2 mm. |
in adaxial tuft, yellow, 0.5–2.5 mm. |
Flowers | inner tepals rose to red-purple, bronze-purple, or yellow, sometimes pale greenish yellow or whitish, spatulate, 18–35 mm, emarginate-apiculate; filaments deep purple to pink-purple, sometimes green; anthers pale yellow; style white or pink to purple distally; stigma lobes white to cream. |
inner tepals greenish basally to reddish bronze, tinged magenta apically, spatulate-apiculate, 15–25 mm; filaments greenish basally to bronze distally; anthers yellow; style greenish bronze; stigma lobes cream. |
Fruits | rarely proliferating, yellow, sometimes tinged reddish to purplish, broadly cylindric, 20–50 × 17–30 mm, fleshy, strongly tuberculate, spineless; areoles 28–50(–62); tubercles longer in distal portion of fruit; umbilicus to 10 mm deep. |
green becoming red, obovoid to cylindric, 13–34 × 10–20 mm, fleshy, low tuberculate to smooth, spineless; tubercles subequal; umbilicus 3–6 mm deep; areoles 16–28. |
Seeds | pale yellow, suborbicular to oval in outline, flattened to warped, 4–5 × 3–4 mm, sides with 0–3 large depressions; girdle smooth. |
tan, squarish to oval, 4–5 × 3.5–4 mm, warped, sides smooth to lumpy; girdle broad, low-ridged. |
2n | = 22. |
= 44. |
Cylindropuntia spinosior |
Cylindropuntia kleiniae |
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Phenology | Flowering spring–early summer (Apr–Aug). | Flowering late spring–early summer (May–Jun). |
Habitat | Desert and plains grasslands, extending onto Sonoran Desert flats, sandy to loamy soils | Chihuahuan Desert, creosote bush-mesquite flats, limestone rocky slopes |
Elevation | 300-2000 m (1000-6600 ft) | 800-1800 m (2600-5900 ft) |
Distribution |
AZ; NM; Mexico (Chihuahua, Sonora)
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NM; TX; Mexico (Chihuahua, Coahuila, Durango, Nuevo León)
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Discussion | Cylindropuntia spinosior forms hybrids with C. acanthocarpa var. major, C. arbuscula, C. fulgida (see 6. C. ×kelvinensis), C. leptocaulis (see discussion under 3. C. ×tetracantha), and C. versicolor (= C. ×grantiorum P. V. Heath). Introgression between C. spinosior and C. imbricata occurs in a more or less continuous band from central New Mexico to Chihuahua, Mexico. The hybrids between C. spinosior and C. acanthocarpa var. major are sprawling shrubs with irregular branching pattern and have spine clusters with one or more spines longer than others and fleshy fruits, some with one to few spines per fruit. Chromosome numbers of hybrids are all reported as 2n = 22. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In western Texas and northern Mexico, Cylindropuntia kleiniae forms hybrids with C. leptocaulis (= C. ×antoniae P. V. Heath) that have varying degrees of morphologic intermediacy and chromosome number of 2n = 33. In northern Mexico, C. kleiniae hybridizes with C. imbricata var. imbricata. Cylindropuntia kleiniae itself may be of hybrid origin. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 4. | FNA vol. 4, p. 107. |
Parent taxa | Cactaceae > subfam. Opuntioideae > Cylindropuntia | Cactaceae > subfam. Opuntioideae > Cylindropuntia |
Sibling taxa | ||
Synonyms | Opuntia whipplei var. spinosior, Opuntia spinosior, Opuntia spinosior var. neomexicana | Opuntia kleiniae, Opuntia wrightii |
Name authority | (Engelmann) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 126. (1935) | (de Candolle) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kactus-ABC, 123. (1935) |
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