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branch pencil cholla, diamond cholla, pencil cactus

wolf cholla, Wolf's cholla, Wolf's opuntia

Habit Trees or shrubs, intricately branched, 0.5–2 m. Stem segments firmly attached, green drying gray and ropelike, cylindric, 2–8(–10) × 0.4–1 cm; tubercles rhombic, convex (flattened upon drying), 0.4–0.8 cm; areoles subcircular abaxially, adaxially becoming usually deltate-linear; glochid-bearing portion protruding distally, wedged between bases of 2 adjacent tubercles, (3–)4–7 × 1–1.5(–2) mm; wool tan to white. Shrubs, ± erect, densely branched, 0.5–1.1(–1.5) m. Stem segments yellow- to gray-green, 6–40 × 2.5–4 cm; tubercles prominent, 1–1.5(–2) cm; areoles subcircular to obdeltate, 5–7 × 4.5–5 mm; wool tan to yellow, aging gray.
Spines

0–5 per areole, usually in distal areoles or sometimes absent or nearly so, tan to red-brown to deep purple, aging gray;

major abaxial spines 0–1(–2), the longest spine spreading, (1.5–)2.5–6 cm;

adaxial spines usually reflexed, short to ± 1 cm;

sheaths baggy.

12–25(–30) per areole, gold to pale brown, aging dark brown, longest to 3 cm;

sheaths translucent yellow to pale brown.

Glochids

in subcircular to linear adaxial tuft, yellow to tan to brown, to 2 mm.

in adaxial tuft and scattered along margin, yellow to pale brown, of irregular lengths.

Flowers

inner tepals bronze-red ± suffused rose, with mid stripes darker, ovate, 6–13 mm, acute-apiculate to attenuate;

filaments greenish;

anthers yellow;

style whitish or blushed with rose-pink or light green;

stigma lobes whitish.

inner tepals yellow-green suffused with bronze, pale red, or purple-brown, 20–35 mm;

filaments dark bronze-red to magenta;

anthers yellow, sometimes sterile;

style rose-pink (at least distally);

stigma lobes cream.

Fruits

maturing tan, ellipsoid to stipitate-ellipsoid, 15–30 × 10–15 mm, dry at maturity, tuberculate, developing increasingly burlike, with many bristlelike spines;

areoles (32–)40–66, evenly spaced, woolly.

grayish tan, 25–30 × 15–30 mm, dry, tuberculate, densely spiny;

tubercles prominent, subequal or basal ones longest;

areoles 24–32.

Seeds

pale yellow to tan-gray, angular to squarish in outline, warped, 4–4.5 × 3.5–4 mm, sides irregularly concave-convex;

girdle smooth.

rare, tan, circular, flattened, 5–6 mm diam., with 1–2 shallow depressions;

girdle smooth.

2n

= 22, 44.

= 66.

Cylindropuntia ramosissima

Cylindropuntia wolfii

Phenology Flowering spring–summer (Apr–Aug). Flowering spring (Mar–May).
Habitat Mojave and Sonoran deserts, washes, flats, and bajadas, sandy loam, desert pavement, stony volcanic substrates Dry, rocky slopes
Elevation 50-1100 m (200-3600 ft) 300-1200 m (1000-3900 ft)
Distribution
from FNA
AZ; CA; NV; Mexico (Baja California, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico (Baja California)
[BONAP county map]
Source FNA vol. 4, p. 118. FNA vol. 4, p. 117.
Parent taxa Cactaceae > subfam. Opuntioideae > Cylindropuntia Cactaceae > subfam. Opuntioideae > Cylindropuntia
Sibling taxa
C. abyssi, C. acanthocarpa, C. arbuscula, C. bigelovii, C. californica, C. davisii, C. echinocarpa, C. fulgida, C. ganderi, C. imbricata, C. kleiniae, C. leptocaulis, C. munzii, C. prolifera, C. spinosior, C. tunicata, C. versicolor, C. whipplei, C. wolfii, C. ×kelvinensis, C. ×tetracantha
C. abyssi, C. acanthocarpa, C. arbuscula, C. bigelovii, C. californica, C. davisii, C. echinocarpa, C. fulgida, C. ganderi, C. imbricata, C. kleiniae, C. leptocaulis, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. versicolor, C. whipplei, C. ×kelvinensis, C. ×tetracantha
Synonyms Opuntia ramosissima, Opuntia tessellata Opuntia echinocarpa var. wolfii, Opuntia wolfii
Name authority (Engelmann) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktu s-ABC, 122. (1935) (L. D. Benson) M. A. Baker: J. Arizona-Nevada Acad. Sci. 33: 150. (2001)
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