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coyonostyle, tree cholla

kelvin pricklypear

Habit Trees, with short trunks, or large shrubs, widely branching, (1–)1.5–2.5(–5) m. Stem segments firmly attached, whorled or subwhorled, gray-green, cylindric to weakly clavate, 10–40 × 1.5–4 cm; tubercles very prominent, widely spaced, (1.5–)2–5 cm; areoles elliptic, 5–8× 3–4 mm; wool yellow to tan, aging gray to black. Trees 1.5–2 m; trunk commonly branched; crown open to compact usually bearing whorled branches.
Stem

segments often easily dislodged, if terminal, green or purple, 5–13 × 1.8–3 cm;

tubercles prominent, broadly oval, 0.7–1.2(–1.5) cm;

areoles obdeltate to rhombic, 4.5–6 × 3–5 mm;

wool yellow, aging gray to black.

Spines

(5–)8–15(–30) per areole or spineless to nearly spineless, usually at most areoles, not obscuring stems, spreading, straight or slightly curved, silver to yellow to usually ± red or tan to brown, stout, terete or sometimes flattened basally, 8–30(–40) mm subequal;

sheaths silver to yellow to usually tan to dirty white, yellow tipped.

0–8(–13) per areole, sometimes with 1–3 bristlelike spines along areole margins at most areoles, usually slightly interlacing with spines of adjacent areoles, yellowish to tan, sometimes with pinkish hue, to red-brown and partially gray coated, aging brown-gray to gray;

major abaxial spines angular-flattened, usually reflexed, the longest to 30 mm;

major adaxial spines subterete, erect-divergent, the longest to 18 mm;

sheaths uniformly whitish or tipped yellow to gold, slightly baggy.

Glochids

in dense adaxial tuft, pale yellow, 0.5–3 mm.

in inconspicuous adaxial crescent, yellow, to 1 mm.

Flowers

inner tepals dark pink to magenta to red-magenta, obovate, apiculate, 15–35 mm;

filaments green basally to pink to magenta distally;

anthers yellow;

style light green basally or pink to red-magenta apically;

stigma lobes green or cream.

inner tepals rose to magenta, spatulate, 18–25(–30) mm, emarginate-apiculate;

filaments deep rose;

anthers yellow;

style white, tinged red-purple distally;

stigma lobes white with hint of green.

Fruits

not proliferating, yellow, obovoid, 24–45 × 20–40 mm, fleshy, tuberculate, spineless;

areoles 18–30;

tubercles nearly equal in length or longer in proximal portion of fruit, prominent (occasionally smooth at maturity);

umbilicus 7–14 mm deep.

usually sterile, sometimes forming short chains, yellow-green to yellow, sometimes purplish, 25–45 × 15–22 mm, fleshy, tuberculate, spineless;

tubercles progressively longer toward fruit apex;

umbilicus 4–7 mm deep;

areoles 32–44.

Seeds

yellow-tan, subcircular to angled, warped, 2.5–4 mm diam., sides smooth to slightly lumpy;

girdle usually narrow, not protruding.

pale yellow, orbicular to angular or squarish in outline, flattened to warped, 3.5–4 × 3–4 mm, sides with 0–2 large depressions;

hilum sometimes pointed;

girdle smooth.

2n

= 22, 33.

Cylindropuntia imbricata

Cylindropuntia ×kelvinensis

Phenology Flowering spring (Apr–Jun).
Habitat Sonoran desert scrub, edges of grasslands, rocky flats and slopes, rolling hills
Elevation 500-1000 m (1600-3300 ft)
Distribution
from FNA
AZ; CO; KS; NM; OK; TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ
[BONAP county map]
Discussion

Varieties 2+ (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants trees 3-5 m; stem segments 12-40 cm; tubercles widely spaced; spines, and sheaths when present, usually tan to dirty white or ± yellow
var. imbricata
1. Plants erect shrubs to 1.2 m; stem segments 10-20 cm; tubercles narrowly spaced; spines and sheaths silvery
var. argentea
Source FNA vol. 4, p. 112. FNA vol. 4, p. 108.
Parent taxa Cactaceae > subfam. Opuntioideae > Cylindropuntia Cactaceae > subfam. Opuntioideae > Cylindropuntia
Sibling taxa
C. abyssi, C. acanthocarpa, C. arbuscula, C. bigelovii, C. californica, C. davisii, C. echinocarpa, C. fulgida, C. ganderi, C. kleiniae, C. leptocaulis, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. versicolor, C. whipplei, C. wolfii, C. ×kelvinensis, C. ×tetracantha
C. abyssi, C. acanthocarpa, C. arbuscula, C. bigelovii, C. californica, C. davisii, C. echinocarpa, C. fulgida, C. ganderi, C. imbricata, C. kleiniae, C. leptocaulis, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. versicolor, C. whipplei, C. wolfii, C. ×tetracantha
Subordinate taxa
C. imbricata var. argentea, C. imbricata var. imbricata
Synonyms family cactaceae Cereus imbricatusH, Opuntia imbricata Opuntia kelvinensis
Name authority (Haworth) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 125. (1935) (V. E. Grant & K. A. Grant) P. V. Heath: Calyx 4: 142. (1994)
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