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coyonostyle, tree cholla

Gander cholla, Gander's buckhorn cholla, Gander's cholla

Habit Trees, with short trunks, or large shrubs, widely branching, (1–)1.5–2.5(–5) m. Stem segments firmly attached, whorled or subwhorled, gray-green, cylindric to weakly clavate, 10–40 × 1.5–4 cm; tubercles very prominent, widely spaced, (1.5–)2–5 cm; areoles elliptic, 5–8× 3–4 mm; wool yellow to tan, aging gray to black. Shrubs, ascending, with strict branches, 0.5–1.5 m. Stem segments firmly attached, green, (5–)10–26 × (1.7–)2.5–4.5 cm; tubercles prominent, narrowly elliptic to oval, often subribbed, (0.9–)1.3–2.6 cm; areoles elliptic to obdeltate, 4–7 × 2.5–5 mm; wool yellowish tan, aging gray.
Spines

(5–)8–15(–30) per areole or spineless to nearly spineless, usually at most areoles, not obscuring stems, spreading, straight or slightly curved, silver to yellow to usually ± red or tan to brown, stout, terete or sometimes flattened basally, 8–30(–40) mm subequal;

sheaths silver to yellow to usually tan to dirty white, yellow tipped.

11–18(–28) per areole, at most areoles, moderately to wholly obscuring stems, spreading or erect, whitish, pale yellow to pinkish tan to reddish, aging brown or black, terete or abaxial spines basally flattened, grading in length, the longest porrect to reflexed, 2–2.5(–3.5) cm, with 0–6 bristlelike spines at areole margins;

sheaths slightly baggy, whitish becoming yellow to golden apically.

Glochids

in dense adaxial tuft, pale yellow, 0.5–3 mm.

in thin adaxial crescent, yellow or tan to rusty, 0.5–6 mm.

Flowers

inner tepals dark pink to magenta to red-magenta, obovate, apiculate, 15–35 mm;

filaments green basally to pink to magenta distally;

anthers yellow;

style light green basally or pink to red-magenta apically;

stigma lobes green or cream.

inner tepals greenish yellow, often with tips reddish abaxially, spatulate, 22–30 mm, apiculate to emarginate-apiculate;

filaments yellow-green to green;

anthers yellow to orange-yellow;

style cream to yellow, sometimes basally greenish;

stigma lobes greenish cream, cream or yellow.

Fruits

not proliferating, yellow, obovoid, 24–45 × 20–40 mm, fleshy, tuberculate, spineless;

areoles 18–30;

tubercles nearly equal in length or longer in proximal portion of fruit, prominent (occasionally smooth at maturity);

umbilicus 7–14 mm deep.

yellowish, drying tan, globose, tuberculate, budlike with spines similar to those of stem segments but slightly shorter, often reflexed;

proximal tubercles slightly elongated if fruits fertile;

umbilicus 4–5 mm deep;

areoles 22–32(–40).

Seeds

yellow-tan, subcircular to angled, warped, 2.5–4 mm diam., sides smooth to slightly lumpy;

girdle usually narrow, not protruding.

pale yellow, angular-suborbicular, flattened to slightly warped, 4.5–6 × 4.5–5.7 mm, sides with 3–5 depressions;

girdle smooth, not protruding.

2n

= 22.

Cylindropuntia imbricata

Cylindropuntia ganderi

Phenology Flowering spring (Mar–May).
Habitat Desert alluvial flats, bajadas, sandy, gravelly to rocky substrates
Elevation 100-1000 m (300-3300 ft)
Distribution
from FNA
AZ; CO; KS; NM; OK; TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2+ (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In the north part of its range, Cylindropuntia ganderi intergrades with C. californica var. parkeri.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants trees 3-5 m; stem segments 12-40 cm; tubercles widely spaced; spines, and sheaths when present, usually tan to dirty white or ± yellow
var. imbricata
1. Plants erect shrubs to 1.2 m; stem segments 10-20 cm; tubercles narrowly spaced; spines and sheaths silvery
var. argentea
Source FNA vol. 4, p. 112. FNA vol. 4, p. 115.
Parent taxa Cactaceae > subfam. Opuntioideae > Cylindropuntia Cactaceae > subfam. Opuntioideae > Cylindropuntia
Sibling taxa
C. abyssi, C. acanthocarpa, C. arbuscula, C. bigelovii, C. californica, C. davisii, C. echinocarpa, C. fulgida, C. ganderi, C. kleiniae, C. leptocaulis, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. versicolor, C. whipplei, C. wolfii, C. ×kelvinensis, C. ×tetracantha
C. abyssi, C. acanthocarpa, C. arbuscula, C. bigelovii, C. californica, C. davisii, C. echinocarpa, C. fulgida, C. imbricata, C. kleiniae, C. leptocaulis, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. versicolor, C. whipplei, C. wolfii, C. ×kelvinensis, C. ×tetracantha
Subordinate taxa
C. imbricata var. argentea, C. imbricata var. imbricata
Synonyms family cactaceae Cereus imbricatusH, Opuntia imbricata Opuntia acanthocarpa subsp. ganderi, Opuntia acanthocarpa var. ganderi, Opuntia ganderi
Name authority (Haworth) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 125. (1935) (C. B. Wolf) Rebman & Pinkava: J. Arizona-Nevada Acad. Sci. 33: 150. (2001)
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