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Gander cholla, Gander's buckhorn cholla, Gander's cholla

Christmas cactus, desert Christmas cactus, desert Christmas cholla, tasajillo

Habit Shrubs, ascending, with strict branches, 0.5–1.5 m. Stem segments firmly attached, green, (5–)10–26 × (1.7–)2.5–4.5 cm; tubercles prominent, narrowly elliptic to oval, often subribbed, (0.9–)1.3–2.6 cm; areoles elliptic to obdeltate, 4–7 × 2.5–5 mm; wool yellowish tan, aging gray. Shrubs or small trees, sparingly to densely branched, 0.5–1.8 m, usually bearing similar, commonly spineless terminal branchlets arranged at right angles along major axes.
Stem

segments usually alternate, gray-green or purplish, 2–8 × 0.3–0.5 cm;

tubercles linear, drying as elongate, riblike wrinkles, 1.1–2(–3) cm;

areoles broadly elliptic, (1–)1.5–3.5 × 0.7–2 mm;

wool white to yellow, aging gray.

Spines

11–18(–28) per areole, at most areoles, moderately to wholly obscuring stems, spreading or erect, whitish, pale yellow to pinkish tan to reddish, aging brown or black, terete or abaxial spines basally flattened, grading in length, the longest porrect to reflexed, 2–2.5(–3.5) cm, with 0–6 bristlelike spines at areole margins;

sheaths slightly baggy, whitish becoming yellow to golden apically.

0–1(–3) per areole, usually in apical areoles to well distributed, erect, flexible, straight or arching upward or downward, red-brown with gray to whitish coat, tips yellow, aging red-brown, terete, angular-flattened basally, the longest (4–)14–45 mm;

sheaths gray to purple-gray with yellow to red-brown tips or yellow throughout.

Glochids

in thin adaxial crescent, yellow or tan to rusty, 0.5–6 mm.

in adaxial tuft or crescent to encircling areole, yellow or reddish brown, 1–3(–5) mm.

Flowers

inner tepals greenish yellow, often with tips reddish abaxially, spatulate, 22–30 mm, apiculate to emarginate-apiculate;

filaments yellow-green to green;

anthers yellow to orange-yellow;

style cream to yellow, sometimes basally greenish;

stigma lobes greenish cream, cream or yellow.

inner tepals pale yellow to greenish yellow, sometimes tipped red, narrowly obovate, 5–8 mm, acute, apiculate;

filaments greenish yellow;

anthers yellow;

style yellow;

stigma lobes greenish yellow.

Fruits

yellowish, drying tan, globose, tuberculate, budlike with spines similar to those of stem segments but slightly shorter, often reflexed;

proximal tubercles slightly elongated if fruits fertile;

umbilicus 4–5 mm deep;

areoles 22–32(–40).

occasionally proliferating, yellow to scarlet (rarely green, sometimes tinged purple, becoming yellow), sometimes stipitate, obovoid, 9–15(–27) × 6–7(–12) mm, fleshy, smooth, spineless;

umbilicus 2–4 mm deep;

areoles 16–20.

Seeds

pale yellow, angular-suborbicular, flattened to slightly warped, 4.5–6 × 4.5–5.7 mm, sides with 3–5 depressions;

girdle smooth, not protruding.

pale yellow, suborbicular to squarish and crenate in outline, warped, 3–4.5 mm diam., sides smooth, each with 1–3 large depressions;

girdle smooth or with very narrow ridge.

2n

= 22.

= 22, 33, 44.

Cylindropuntia ganderi

Cylindropuntia leptocaulis

Phenology Flowering spring (Mar–May). Flowering spring–early summer, sometimes fall (Mar–Aug, Oct).
Habitat Desert alluvial flats, bajadas, sandy, gravelly to rocky substrates Deserts, grasslands, chaparrals, oak-juniper woodlands, flats, bajadas and slopes, sandy, loamy to gravelly substrates
Elevation 100-1000 m (300-3300 ft) 40-1500 m (100-4900 ft)
Distribution
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; OK; TX; Mexico (Chihuahua, Coahuila, Durango, Nuevo León, Sonora, Tamaulipas, Zacatecas)
[WildflowerSearch map]
[BONAP county map]
Discussion

In the north part of its range, Cylindropuntia ganderi intergrades with C. californica var. parkeri.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cylindropuntia leptocaulis forms hybrids with C. acanthocarpa var. major (see 3. C. ×tetracantha), C. arbuscula, C. fulgida, C. kleiniae, C. spinosior, C. versicolor (see discussion under C. ×tetracantha), and C. whipplei. Hybrids in central Arizona have flowers intermediate in size to the parents, narrow tuberculate stems bearing 0–1(–2) major spines per areole, and tuberculate, spineless, orange to red fruits. The chromosome number reported for hybrids is 2n = 22.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 4, p. 115. FNA vol. 4, p. 105.
Parent taxa Cactaceae > subfam. Opuntioideae > Cylindropuntia Cactaceae > subfam. Opuntioideae > Cylindropuntia
Sibling taxa
C. abyssi, C. acanthocarpa, C. arbuscula, C. bigelovii, C. californica, C. davisii, C. echinocarpa, C. fulgida, C. imbricata, C. kleiniae, C. leptocaulis, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. versicolor, C. whipplei, C. wolfii, C. ×kelvinensis, C. ×tetracantha
C. abyssi, C. acanthocarpa, C. arbuscula, C. bigelovii, C. californica, C. davisii, C. echinocarpa, C. fulgida, C. ganderi, C. imbricata, C. kleiniae, C. munzii, C. prolifera, C. ramosissima, C. spinosior, C. tunicata, C. versicolor, C. whipplei, C. wolfii, C. ×kelvinensis, C. ×tetracantha
Synonyms Opuntia acanthocarpa subsp. ganderi, Opuntia acanthocarpa var. ganderi, Opuntia ganderi Opuntia leptocaulis, C. leptocaulis var. brevispina, C. leptocaulis var. longispina, Opuntia fragilis var. frutescens, Opuntia frutescens, Opuntia frutescens var. brevispina, Opuntia frutescens var. longispina, Opuntia leptocaulis, Opuntia leptocaulis var. longispina, Opuntia leptocaulis var. vaginata, Opuntia vaginata
Name authority (C. B. Wolf) Rebman & Pinkava: J. Arizona-Nevada Acad. Sci. 33: 150. (2001) (de Candolle) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 122. (1935)
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