Cylindropuntia arbuscula |
Cylindropuntia ramosissima |
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Arizona pencil cholla, pencil cholla |
branch pencil cholla, diamond cholla, pencil cactus |
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Habit | Trees or shrubs, densely branched, 0.5–3 m. Stem segments green, sometimes purple tinged, 6–10.5 × 0.5–1.3 cm; tubercles narrowly elongate, appearing as wrinkles when dry, 1–2 cm; areoles circular, 2–3 mm in diam.; wool tan to brown, aging gray to black. | Trees or shrubs, intricately branched, 0.5–2 m. Stem segments firmly attached, green drying gray and ropelike, cylindric, 2–8(–10) × 0.4–1 cm; tubercles rhombic, convex (flattened upon drying), 0.4–0.8 cm; areoles subcircular abaxially, adaxially becoming usually deltate-linear; glochid-bearing portion protruding distally, wedged between bases of 2 adjacent tubercles, (3–)4–7 × 1–1.5(–2) mm; wool tan to white. |
Spines | 0–2(–3) per areole, sparsely distributed along stem, usually deflexed, pale yellow or red-brown aging black, stout, the longest 0.8–3.5(–5) cm; sheaths loose fitting, yellowish brown. |
0–5 per areole, usually in distal areoles or sometimes absent or nearly so, tan to red-brown to deep purple, aging gray; major abaxial spines 0–1(–2), the longest spine spreading, (1.5–)2.5–6 cm; adaxial spines usually reflexed, short to ± 1 cm; sheaths baggy. |
Glochids | in adaxial tuft and marginal, encircling areole, pale yellow. |
in subcircular to linear adaxial tuft, yellow to tan to brown, to 2 mm. |
Flowers | inner tepals green- or orange-bronze, spatulate, 17–20 mm, apiculate; filaments dark green-bronze; anthers yellow; style whitish basally to light orange apically; stigma lobes very pale green. |
inner tepals bronze-red ± suffused rose, with mid stripes darker, ovate, 6–13 mm, acute-apiculate to attenuate; filaments greenish; anthers yellow; style whitish or blushed with rose-pink or light green; stigma lobes whitish. |
Fruits | green, becoming yellowish apically, sometimes tinged red to purplish at areoles, commonly sterile, then narrow, tuberculate, to 2.5 cm, fertile ones stipitate, obconic, 20–50 × 15–35 mm, fleshy, becoming smooth, spineless; umbilicus 3–4 mm deep; areoles 15–17(–34). |
maturing tan, ellipsoid to stipitate-ellipsoid, 15–30 × 10–15 mm, dry at maturity, tuberculate, developing increasingly burlike, with many bristlelike spines; areoles (32–)40–66, evenly spaced, woolly. |
Seeds | pale yellow, angularly circular to oblong, thick, 3–5 mm diam., lumpy; girdle smooth, broad, not protruding. |
pale yellow to tan-gray, angular to squarish in outline, warped, 4–4.5 × 3.5–4 mm, sides irregularly concave-convex; girdle smooth. |
2n | = 66. |
= 22, 44. |
Cylindropuntia arbuscula |
Cylindropuntia ramosissima |
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Phenology | Flowering spring (Apr–Jun). | Flowering spring–summer (Apr–Aug). |
Habitat | Sonoran Desert flats, bajadas, desert scrub | Mojave and Sonoran deserts, washes, flats, and bajadas, sandy loam, desert pavement, stony volcanic substrates |
Elevation | 300-1000 m (1000-3300 ft) | 50-1100 m (200-3600 ft) |
Distribution |
AZ; Mexico (Sonora)
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AZ; CA; NV; Mexico (Baja California, Sonora)
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Discussion | Cylindropuntia arbuscula forms hybrids with C. leptocaulis in south-central Arizona; those hybrids have narrow, obscurely tuberculate stems and reddish fruits nearly the size of those of C. arbuscula and chromosome number of 2n = 55. Hybrids between C. arbuscula and C. spinosior [= C. ×neoarbuscula (Griffiths) F. M. Knuth] have large green fruits, which often split open, and on stem segments distal areoles that usually bear three or four spines to 2.5 cm. Hybrids between C. arbuscula and C. versicolor [= C. ×vivipara (Rose) F. M. Knuth] have large green fruits, which often split open, and distal stem segment areoles that bear one or two spines to 1.5 cm. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 4, p. 106. | FNA vol. 4, p. 118. |
Parent taxa | Cactaceae > subfam. Opuntioideae > Cylindropuntia | Cactaceae > subfam. Opuntioideae > Cylindropuntia |
Sibling taxa | ||
Synonyms | Opuntia arbuscula | Opuntia ramosissima, Opuntia tessellata |
Name authority | (Engelmann) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktus-ABC, 123. (1935) | (Engelmann) F. M. Knuth: in C. Backeberg and F. M. Knuth, Kaktu s-ABC, 122. (1935) |
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