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bush-clover dodder, field dodder, five-angle dodder, western field dodder

Jepson's dodder

Stems

yellow to orange, slender to medium.

pale yellow, slender.

Inflorescences

dense, corymbiform to glomerulate;

bracts at base of clusters 1, at base of pedicels 0 or 1, ovate or ovate-triangular to lanceolate, membranous, margins entire, apex acute.

moderately dense, corymbiform or glomerulate;

bracts at base of clusters 1, at base of pedicels 0 or 1, ovate to lanceolate, membranous, margins entire, apex acute.

Pedicels

0.5–3(–4.5) mm.

0.5–1.5 mm, papillate.

Flowers

(4 or)5-merous, 1.4–2.5 mm, membranous, corolla lobes sometimes papillate;

calyx yellow to brown, angled, cupulate, equaling corolla tube length, divided 1/2–2/3 its length, ± reticulate, shiny or not, lobes broadly ovate to rhombic, base auriculate, overlapping, forming prominent angles at sinuses, margins entire, midvein not carinate, without multicellular protuberances, apex rounded;

corolla whitish, drying yellow to brown, 1.2–2.2 mm, tube campanulate, 0.7–1.2 mm, not saccate, lobes spreading, triangular-lanceolate, equaling corolla tube length, margins entire, apex acute to acuminate, inflexed;

infrastaminal scales ovate to oblong, 0.7–1.4 mm, equaling or longer than corolla tube length, bridged at 0.3–0.5 mm, rounded, ± uniformly densely fimbriate, 0.15–0.25 mm;

stamens exserted, shorter than corolla lobes;

filaments 0.3–0.4 mm;

anthers 0.2–0.3 × 0.2–0.3 mm;

styles filiform, 0.7–1.1 mm, equaling ovary.

5-merous, 2–3 mm, fleshy, perianth cells convex, domelike, perianth papillate;

calyx yellow-brownish, shallowly cupulate, 1/2 corolla tube length, divided 1/2 its length, not reticulate or shiny, lobes triangular, bases not overlapping, margins entire, midvein not carinate, apex acute;

corolla white, drying brownish, 1.8–2.8 mm, tube campanulate-globulose, 1.3–2 mm, later urceolate, not saccate, lobes erect, ovate-triangular, 1/3–1/2 corolla tube length, margins entire, apex acute, inflexed;

infrastaminal scales absent;

stamens included, shorter than corolla lobes;

filaments 0.2–0.3 mm;

anthers 0.2–0.3 × 0.2–0.3 mm;

styles subulate, 0.4–0.8 mm, much shorter than ovary.

Capsules

depressed-globose to ovoid, 1.9–2.4 × 1.6–2.5 mm, not thickened or raised around relatively medium-sized to large interstylar aperture, translucent or not, base ± enveloped by withered corolla, indehiscent.

subglobose or globose to slightly depressed-globose, 2–3 × 2–3.5 mm, thickened and raised around relatively large interstylar aperture, translucent, surrounded by withered corolla, indehiscent.

Seeds

4, angled, subglobose to broadly ellipsoid, 0.9–1.1 × 0.8–1 mm, hilum region subterminal.

2–4, broadly ellipsoid to subglobose, obcompressed, 0.9–1.1 × 0.8–1 mm, hilum subterminal.

2n

= 56.

Cuscuta pentagona

Cuscuta jepsonii

Phenology Flowering Apr–Nov. Flowering Jul–Sep.
Habitat Hosts: herbs. mixed forests of Pinus jeffreyi, P. ponderosa, and Pseudotsuga menziesii.
Elevation 0–900 m. (0–3000 ft.) 1100–1600 m. (3600–5200 ft.)
Distribution
from FNA
AL; AR; DC; DE; FL; GA; IL; IN; KS; MA; MD; MI; MN; MO; MS; MT; NC; ND; NJ; NY; OK; PA; SC; SD; TN; TX; VA; MB
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Cuscuta pentagona apparently has not spread outside of North America, where it is less common than C. campestris. It is currently included in North American noxious weeds lists although there is no evidence it attacks crops.

G. Engelmann (1859) distinguished four varieties of Cuscuta pentagona: var. calycina, var. microcalyx, var. pentagona, and var. verrucosa (Engelmann) Yuncker. T. G. Yuncker (1932, 1965) treated var. calycina and var. verrucosa at specific rank and provided a new name for each: C. campestris and C. glabrior, respectively. Cuscuta glabrior is currently accepted by all the North American overviews; C. campestris has been persistently considered a synonym of C. pentagona despite morphological and evolutionary evidence that the two are distinct (M. Costea et al. 2006c, 2015).

Cuscuta pentagona differs from C. campestris in its rhombic to ovate, auriculate calyx lobes with overlapping bases that form angles at sinuses and in its smaller flowers, capsules, and seeds.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cuscuta jepsonii belongs to sect. Californicae Yunker (M. Costea and S. Stefanović 2009) and was presumed extinct until collected again from Yosemite National Park (Mariposa County) in 2009.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 14. FNA vol. 14.
Parent taxa Convolvulaceae > Cuscuta > subg. Grammica Convolvulaceae > Cuscuta > subg. Grammica
Sibling taxa
C. americana, C. approximata, C. azteca, C. boldinghii, C. brachycalyx, C. californica, C. campestris, C. cephalanthi, C. chinensis, C. compacta, C. coryli, C. cuspidata, C. dentatasquamata, C. denticulata, C. draconella, C. epilinum, C. epithymum, C. erosa, C. europaea, C. exaltata, C. glabrior, C. glomerata, C. gronovii, C. harperi, C. howelliana, C. indecora, C. japonica, C. jepsonii, C. legitima, C. leptantha, C. liliputana, C. mitriformis, C. nevadensis, C. obtusiflora, C. occidentalis, C. odontolepis, C. pacifica, C. plattensis, C. polygonorum, C. rostrata, C. runyonii, C. salina, C. squamata, C. suaveolens, C. subinclusa, C. suksdorfii, C. tuberculata, C. umbellata, C. umbrosa, C. warneri
C. americana, C. approximata, C. azteca, C. boldinghii, C. brachycalyx, C. californica, C. campestris, C. cephalanthi, C. chinensis, C. compacta, C. coryli, C. cuspidata, C. dentatasquamata, C. denticulata, C. draconella, C. epilinum, C. epithymum, C. erosa, C. europaea, C. exaltata, C. glabrior, C. glomerata, C. gronovii, C. harperi, C. howelliana, C. indecora, C. japonica, C. legitima, C. leptantha, C. liliputana, C. mitriformis, C. nevadensis, C. obtusiflora, C. occidentalis, C. odontolepis, C. pacifica, C. pentagona, C. plattensis, C. polygonorum, C. rostrata, C. runyonii, C. salina, C. squamata, C. suaveolens, C. subinclusa, C. suksdorfii, C. tuberculata, C. umbellata, C. umbrosa, C. warneri
Synonyms C. arvensis, C. pentagona var. microcalyx, Grammica pentagona
Name authority Engelmann: Amer. J. Sci. Arts 43: 340, plate 6, figs. 22–24. (1842) Yuncker: Illinois Biol. Monogr. 6: 149, plate 9, fig. 52. (1921)
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