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bush-clover dodder, field dodder, five-angle dodder, western field dodder

Harper's dodder

Stems

yellow to orange, slender to medium.

orange-yellow, filiform.

Inflorescences

dense, corymbiform to glomerulate;

bracts at base of clusters 1, at base of pedicels 0 or 1, ovate or ovate-triangular to lanceolate, membranous, margins entire, apex acute.

loose, corymbiform;

bracts at base of clusters 1, at base of pedicels 0(or 1), ovate-triangular, membranous, margins entire, apex acute.

Pedicels

0.5–3(–4.5) mm.

0.5–2.5(–3) mm.

Flowers

(4 or)5-merous, 1.4–2.5 mm, membranous, corolla lobes sometimes papillate;

calyx yellow to brown, angled, cupulate, equaling corolla tube length, divided 1/2–2/3 its length, ± reticulate, shiny or not, lobes broadly ovate to rhombic, base auriculate, overlapping, forming prominent angles at sinuses, margins entire, midvein not carinate, without multicellular protuberances, apex rounded;

corolla whitish, drying yellow to brown, 1.2–2.2 mm, tube campanulate, 0.7–1.2 mm, not saccate, lobes spreading, triangular-lanceolate, equaling corolla tube length, margins entire, apex acute to acuminate, inflexed;

infrastaminal scales ovate to oblong, 0.7–1.4 mm, equaling or longer than corolla tube length, bridged at 0.3–0.5 mm, rounded, ± uniformly densely fimbriate, 0.15–0.25 mm;

stamens exserted, shorter than corolla lobes;

filaments 0.3–0.4 mm;

anthers 0.2–0.3 × 0.2–0.3 mm;

styles filiform, 0.7–1.1 mm, equaling ovary.

4(or 5)-merous, 0.9–1.1(–1.5) mm, fleshy, corolla lobes papillate;

calyx brownish yellow, angled, cupulate, equaling corolla tube length, divided 1/2–2/3 its length, ± reticulate, rarely shiny, lobes broadly ovate-rhombic, bases ± auriculate, overlapping, forming prominent angles at sinuses, margins entire, midvein ± carinate, apex rounded;

corolla white, drying cream to brownish, 0.9–1.2 mm, tube campanulate, 0.5–0.7 mm, not saccate, lobes reflexed, triangular-ovate, equaling corolla tube length, margins entire, apex subacute to acute, inflexed;

infrastaminal scales narrowly oblong to obovate, 0.6–0.9 mm, equaling or longer than corolla tube length, bridged at 0.2–0.3 mm, rounded, sparsely fimbriate, fimbriae 0.1–0.3 mm;

stamens exserted, slightly shorter than corolla lobes;

filaments 0.1–0.2 mm;

anthers 0.2–0.3 × 0.2–0.3 mm;

styles filiform, 0.5–0.9 mm, shorter than ovary.

Capsules

depressed-globose to ovoid, 1.9–2.4 × 1.6–2.5 mm, not thickened or raised around relatively medium-sized to large interstylar aperture, translucent or not, base ± enveloped by withered corolla, indehiscent.

globose to ovoid, 1.2–2.3 × 1.2–1.6 mm, not thickened around relatively small to moderately large interstylar aperture, ± translucent, withered corolla enveloping 1/4–1/3 of base, indehiscent.

Seeds

4, angled, subglobose to broadly ellipsoid, 0.9–1.1 × 0.8–1 mm, hilum region subterminal.

1 or 2, subglobose, with longitudinal groove on adaxial face, 0.9–1.1 × 0.8–1.1 mm, hilum region terminal.

2n

= 56.

Cuscuta pentagona

Cuscuta harperi

Phenology Flowering Apr–Nov. Flowering Sep–Nov.
Habitat Hosts: herbs. sandstone, less frequently, granite outcrops.
Elevation 0–900 m. (0–3000 ft.) 200–600 m. (700–2000 ft.)
Distribution
from FNA
AL; AR; DC; DE; FL; GA; IL; IN; KS; MA; MD; MI; MN; MO; MS; MT; NC; ND; NJ; NY; OK; PA; SC; SD; TN; TX; VA; MB
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; GA
[BONAP county map]
Discussion

Cuscuta pentagona apparently has not spread outside of North America, where it is less common than C. campestris. It is currently included in North American noxious weeds lists although there is no evidence it attacks crops.

G. Engelmann (1859) distinguished four varieties of Cuscuta pentagona: var. calycina, var. microcalyx, var. pentagona, and var. verrucosa (Engelmann) Yuncker. T. G. Yuncker (1932, 1965) treated var. calycina and var. verrucosa at specific rank and provided a new name for each: C. campestris and C. glabrior, respectively. Cuscuta glabrior is currently accepted by all the North American overviews; C. campestris has been persistently considered a synonym of C. pentagona despite morphological and evolutionary evidence that the two are distinct (M. Costea et al. 2006c, 2015).

Cuscuta pentagona differs from C. campestris in its rhombic to ovate, auriculate calyx lobes with overlapping bases that form angles at sinuses and in its smaller flowers, capsules, and seeds.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cuscuta harperi is closely allied with C. pentagona (M. Costea et al. 2015), from which it differs in smaller, four-merous flowers.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 14. FNA vol. 14.
Parent taxa Convolvulaceae > Cuscuta > subg. Grammica Convolvulaceae > Cuscuta > subg. Grammica
Sibling taxa
C. americana, C. approximata, C. azteca, C. boldinghii, C. brachycalyx, C. californica, C. campestris, C. cephalanthi, C. chinensis, C. compacta, C. coryli, C. cuspidata, C. dentatasquamata, C. denticulata, C. draconella, C. epilinum, C. epithymum, C. erosa, C. europaea, C. exaltata, C. glabrior, C. glomerata, C. gronovii, C. harperi, C. howelliana, C. indecora, C. japonica, C. jepsonii, C. legitima, C. leptantha, C. liliputana, C. mitriformis, C. nevadensis, C. obtusiflora, C. occidentalis, C. odontolepis, C. pacifica, C. plattensis, C. polygonorum, C. rostrata, C. runyonii, C. salina, C. squamata, C. suaveolens, C. subinclusa, C. suksdorfii, C. tuberculata, C. umbellata, C. umbrosa, C. warneri
C. americana, C. approximata, C. azteca, C. boldinghii, C. brachycalyx, C. californica, C. campestris, C. cephalanthi, C. chinensis, C. compacta, C. coryli, C. cuspidata, C. dentatasquamata, C. denticulata, C. draconella, C. epilinum, C. epithymum, C. erosa, C. europaea, C. exaltata, C. glabrior, C. glomerata, C. gronovii, C. howelliana, C. indecora, C. japonica, C. jepsonii, C. legitima, C. leptantha, C. liliputana, C. mitriformis, C. nevadensis, C. obtusiflora, C. occidentalis, C. odontolepis, C. pacifica, C. pentagona, C. plattensis, C. polygonorum, C. rostrata, C. runyonii, C. salina, C. squamata, C. suaveolens, C. subinclusa, C. suksdorfii, C. tuberculata, C. umbellata, C. umbrosa, C. warneri
Synonyms C. arvensis, C. pentagona var. microcalyx, Grammica pentagona
Name authority Engelmann: Amer. J. Sci. Arts 43: 340, plate 6, figs. 22–24. (1842) Small: Fl. S.E. U.S. ed. 2, 1361, 1375. (1913)
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