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bush-clover dodder, field dodder, five-angle dodder, western field dodder

Sonoran dodder

Stems

yellow to orange, slender to medium.

white to creamy yellow, medium.

Inflorescences

dense, corymbiform to glomerulate;

bracts at base of clusters 1, at base of pedicels 0 or 1, ovate or ovate-triangular to lanceolate, membranous, margins entire, apex acute.

loose to moderately dense, paniculiform or corymbiform;

bracts at base of clusters 1, at base of pedicels 0 or 1, ovate-triangular to lanceolate, membranous, margins entire or serrulate-denticulate, apex obtuse to acute.

Pedicels

0.5–3(–4.5) mm.

1.5–6 mm.

Flowers

(4 or)5-merous, 1.4–2.5 mm, membranous, corolla lobes sometimes papillate;

calyx yellow to brown, angled, cupulate, equaling corolla tube length, divided 1/2–2/3 its length, ± reticulate, shiny or not, lobes broadly ovate to rhombic, base auriculate, overlapping, forming prominent angles at sinuses, margins entire, midvein not carinate, without multicellular protuberances, apex rounded;

corolla whitish, drying yellow to brown, 1.2–2.2 mm, tube campanulate, 0.7–1.2 mm, not saccate, lobes spreading, triangular-lanceolate, equaling corolla tube length, margins entire, apex acute to acuminate, inflexed;

infrastaminal scales ovate to oblong, 0.7–1.4 mm, equaling or longer than corolla tube length, bridged at 0.3–0.5 mm, rounded, ± uniformly densely fimbriate, 0.15–0.25 mm;

stamens exserted, shorter than corolla lobes;

filaments 0.3–0.4 mm;

anthers 0.2–0.3 × 0.2–0.3 mm;

styles filiform, 0.7–1.1 mm, equaling ovary.

5-merous, 3.5–4.5 mm, membranous, not papillate;

calyx yellow, cupulate, 3/4 to ± equaling corolla tube length, divided 1/2–2/3 its length, finely reticulate, shiny, lobes orbiculate to oblong-obovate, bases overlapping, margins minutely erose or denticulate, membranous, midvein carinate, apex ± truncate, each not exceeded by a hornlike appendage, 0.1–0.2 mm;

corolla creamy white, drying creamy white to reddish brown, 3.2–4 mm, tube campanulate, 1.5–2.2 mm, not saccate, sometimes with horizontal ridges between stamen attachments, lobes erect or spreading to reflexed, oblong-obovate to orbiculate, shorter than to equaling corolla tube length, margins minutely erose or denticulate, membranous, midvein unevenly carinate, apex ± truncate, each with a hornlike appendage 0.1–0.2 mm, not exceeding apex, straight;

infrastaminal scales oblong to ± truncate, 1.5–2 mm, 3/4 to equaling corolla tube length, bridged at 0.4–0.6 mm, uniformly densely fimbriate, fimbriae 0.2–0.4 mm;

stamens exserted, shorter than corolla lobes;

filaments 0.4–1 mm;

anthers 0.7–1 × 0.4–0.5 mm;

styles stout, subulate, 1.8–3.2 mm, longer than ovary.

Capsules

depressed-globose to ovoid, 1.9–2.4 × 1.6–2.5 mm, not thickened or raised around relatively medium-sized to large interstylar aperture, translucent or not, base ± enveloped by withered corolla, indehiscent.

globose, 2–2.5 × 2.2–2.5 mm, thickened, not raised around inconspicuous interstylar aperture, not translucent, withered corolla near middle or apical, dehiscence circumscissile.

Seeds

4, angled, subglobose to broadly ellipsoid, 0.9–1.1 × 0.8–1 mm, hilum region subterminal.

1–4, angled, subglobose to ovoid, 0.9–1.4 × 0.8–1.3 mm, papillate, hilum region terminal.

2n

= 56.

Cuscuta pentagona

Cuscuta erosa

Phenology Flowering Apr–Nov. Flowering Aug–Oct.
Habitat Hosts: herbs. desert scrub.
Elevation 0–900 m. (0–3000 ft.) 400–1300 m. (1300–4300 ft.)
Distribution
from FNA
AL; AR; DC; DE; FL; GA; IL; IN; KS; MA; MD; MI; MN; MO; MS; MT; NC; ND; NJ; NY; OK; PA; SC; SD; TN; TX; VA; MB
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; Mexico (Baja California, Sinaloa, Sonora)
[BONAP county map]
Discussion

Cuscuta pentagona apparently has not spread outside of North America, where it is less common than C. campestris. It is currently included in North American noxious weeds lists although there is no evidence it attacks crops.

G. Engelmann (1859) distinguished four varieties of Cuscuta pentagona: var. calycina, var. microcalyx, var. pentagona, and var. verrucosa (Engelmann) Yuncker. T. G. Yuncker (1932, 1965) treated var. calycina and var. verrucosa at specific rank and provided a new name for each: C. campestris and C. glabrior, respectively. Cuscuta glabrior is currently accepted by all the North American overviews; C. campestris has been persistently considered a synonym of C. pentagona despite morphological and evolutionary evidence that the two are distinct (M. Costea et al. 2006c, 2015).

Cuscuta pentagona differs from C. campestris in its rhombic to ovate, auriculate calyx lobes with overlapping bases that form angles at sinuses and in its smaller flowers, capsules, and seeds.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 14. FNA vol. 14.
Parent taxa Convolvulaceae > Cuscuta > subg. Grammica Convolvulaceae > Cuscuta > subg. Grammica
Sibling taxa
C. americana, C. approximata, C. azteca, C. boldinghii, C. brachycalyx, C. californica, C. campestris, C. cephalanthi, C. chinensis, C. compacta, C. coryli, C. cuspidata, C. dentatasquamata, C. denticulata, C. draconella, C. epilinum, C. epithymum, C. erosa, C. europaea, C. exaltata, C. glabrior, C. glomerata, C. gronovii, C. harperi, C. howelliana, C. indecora, C. japonica, C. jepsonii, C. legitima, C. leptantha, C. liliputana, C. mitriformis, C. nevadensis, C. obtusiflora, C. occidentalis, C. odontolepis, C. pacifica, C. plattensis, C. polygonorum, C. rostrata, C. runyonii, C. salina, C. squamata, C. suaveolens, C. subinclusa, C. suksdorfii, C. tuberculata, C. umbellata, C. umbrosa, C. warneri
C. americana, C. approximata, C. azteca, C. boldinghii, C. brachycalyx, C. californica, C. campestris, C. cephalanthi, C. chinensis, C. compacta, C. coryli, C. cuspidata, C. dentatasquamata, C. denticulata, C. draconella, C. epilinum, C. epithymum, C. europaea, C. exaltata, C. glabrior, C. glomerata, C. gronovii, C. harperi, C. howelliana, C. indecora, C. japonica, C. jepsonii, C. legitima, C. leptantha, C. liliputana, C. mitriformis, C. nevadensis, C. obtusiflora, C. occidentalis, C. odontolepis, C. pacifica, C. pentagona, C. plattensis, C. polygonorum, C. rostrata, C. runyonii, C. salina, C. squamata, C. suaveolens, C. subinclusa, C. suksdorfii, C. tuberculata, C. umbellata, C. umbrosa, C. warneri
Synonyms C. arvensis, C. pentagona var. microcalyx, Grammica pentagona
Name authority Engelmann: Amer. J. Sci. Arts 43: 340, plate 6, figs. 22–24. (1842) Yuncker: Illinois Biol. Monogr. 6: 116, plate 2, fig. 8, plate 10, fig. 61. (1921)
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