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bush-clover dodder, field dodder, five-angle dodder, western field dodder

cuscute des champs, field dodder, five-angle dodder, large-seed alfalfa dodder

Stems

yellow to orange, slender to medium.

yellow to orange, medium.

Inflorescences

dense, corymbiform to glomerulate;

bracts at base of clusters 1, at base of pedicels 0 or 1, ovate or ovate-triangular to lanceolate, membranous, margins entire, apex acute.

dense, corymbiform or glomerulate;

bracts at base of clusters 1, at base of pedicels 0 or 1, ovate or ovate-triangular to lanceolate, membranous, margins entire, apex acute.

Pedicels

0.5–3(–4.5) mm.

0.3–2.5(–3.5) mm.

Flowers

(4 or)5-merous, 1.4–2.5 mm, membranous, corolla lobes sometimes papillate;

calyx yellow to brown, angled, cupulate, equaling corolla tube length, divided 1/2–2/3 its length, ± reticulate, shiny or not, lobes broadly ovate to rhombic, base auriculate, overlapping, forming prominent angles at sinuses, margins entire, midvein not carinate, without multicellular protuberances, apex rounded;

corolla whitish, drying yellow to brown, 1.2–2.2 mm, tube campanulate, 0.7–1.2 mm, not saccate, lobes spreading, triangular-lanceolate, equaling corolla tube length, margins entire, apex acute to acuminate, inflexed;

infrastaminal scales ovate to oblong, 0.7–1.4 mm, equaling or longer than corolla tube length, bridged at 0.3–0.5 mm, rounded, ± uniformly densely fimbriate, 0.15–0.25 mm;

stamens exserted, shorter than corolla lobes;

filaments 0.3–0.4 mm;

anthers 0.2–0.3 × 0.2–0.3 mm;

styles filiform, 0.7–1.1 mm, equaling ovary.

(4 or)5-merous, 1.9–3.6 mm, membranous, not papillate;

calyx yellow, cupulate, equaling corolla tube length, divided 2/5–3/5 its length, reticulate, shiny, lobes ovate-triangular, bases overlapping, margins entire, midvein not carinate, without multicellular protuberances, apex obtuse to rounded;

corolla creamy white, drying creamy or golden yellow, 2–3.5 mm, tube campanulate, (1.1–)1.5–1.9 mm, not saccate, lobes spreading, triangular to triangular-lanceolate, equaling corolla tube length, margins entire, apex acute to acuminate, inflexed;

infrastaminal scales oblong-ovate to spatulate, rounded, 1.5–2 mm, equaling or exceeding corolla tube length, bridged at 0.3–0.5 mm, uniformly densely fimbriate, fimbriae 0.3–0.4(–0.5) mm;

stamens exserted, shorter than corolla lobes;

filaments 0.4–0.7 mm;

anthers (0.3–)0.4–0.5 × 0.2–0.3 mm;

styles filiform, 0.5–1.6 mm, shorter than to equaling ovary.

Capsules

depressed-globose to ovoid, 1.9–2.4 × 1.6–2.5 mm, not thickened or raised around relatively medium-sized to large interstylar aperture, translucent or not, base ± enveloped by withered corolla, indehiscent.

depressed-globose to depressed, 1.3–2.8 × 1.9–3.8 mm, not thickened or raised around relatively large interstylar aperture, sometimes translucent, to 1/3 enveloped by withered corolla, indehiscent.

Seeds

4, angled, subglobose to broadly ellipsoid, 0.9–1.1 × 0.8–1 mm, hilum region subterminal.

4, angled, subglobose to broadly ellipsoid, 1.1–1.5 × 0.9–1.1 mm, hilum region subterminal.

2n

= 56.

= 56.

Cuscuta pentagona

Cuscuta campestris

Phenology Flowering Apr–Nov. Flowering May–Nov.
Habitat Hosts: herbs. Hosts: Acanthaceae, Astera­ceae, Brassicaceae, Chenopodiaceae, Convolvulaceae, Euphor­biaceae, Fabaceae, Hydrophyllaceae, Polygona­ceae, Solanaceae, Urticaceae, Verbenaceae, and others (M. Costea and F. J. Tardif 2006).
Elevation 0–900 m. (0–3000 ft.) 0–2000 m. (0–6600 ft.)
Distribution
from FNA
AL; AR; DC; DE; FL; GA; IL; IN; KS; MA; MD; MI; MN; MO; MS; MT; NC; ND; NJ; NY; OK; PA; SC; SD; TN; TX; VA; MB
[WildflowerSearch map]
[BONAP county map]
from FNA
AR; AZ; CA; CO; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MS; MT; NC; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; SD; TN; TX; UT; VA; WA; WI; WV; WY; AB; BC; MB; NS; ON; QC; SK; Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Cuscuta pentagona apparently has not spread outside of North America, where it is less common than C. campestris. It is currently included in North American noxious weeds lists although there is no evidence it attacks crops.

G. Engelmann (1859) distinguished four varieties of Cuscuta pentagona: var. calycina, var. microcalyx, var. pentagona, and var. verrucosa (Engelmann) Yuncker. T. G. Yuncker (1932, 1965) treated var. calycina and var. verrucosa at specific rank and provided a new name for each: C. campestris and C. glabrior, respectively. Cuscuta glabrior is currently accepted by all the North American overviews; C. campestris has been persistently considered a synonym of C. pentagona despite morphological and evolutionary evidence that the two are distinct (M. Costea et al. 2006c, 2015).

Cuscuta pentagona differs from C. campestris in its rhombic to ovate, auriculate calyx lobes with overlapping bases that form angles at sinuses and in its smaller flowers, capsules, and seeds.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Reports of Cuscuta campestris from New Brunswick, Newfoundland, and Prince Edward Island have not been verified.

Cuscuta campestris is the most widespread species of the genus in North America and perhaps the most successful and prevalent Cuscuta weed species worldwide; it has been recorded from South America, Europe, Asia, Africa, and Australia. It has been often referred to in North America as C. pentagona, which has smaller flowers and angled calyces. The two species are closely related; C. campestris is a hybrid species and C. pentagona is one of its progenitors (M. Costea et al. 2015b).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 14. FNA vol. 14.
Parent taxa Convolvulaceae > Cuscuta > subg. Grammica Convolvulaceae > Cuscuta > subg. Grammica
Sibling taxa
C. americana, C. approximata, C. azteca, C. boldinghii, C. brachycalyx, C. californica, C. campestris, C. cephalanthi, C. chinensis, C. compacta, C. coryli, C. cuspidata, C. dentatasquamata, C. denticulata, C. draconella, C. epilinum, C. epithymum, C. erosa, C. europaea, C. exaltata, C. glabrior, C. glomerata, C. gronovii, C. harperi, C. howelliana, C. indecora, C. japonica, C. jepsonii, C. legitima, C. leptantha, C. liliputana, C. mitriformis, C. nevadensis, C. obtusiflora, C. occidentalis, C. odontolepis, C. pacifica, C. plattensis, C. polygonorum, C. rostrata, C. runyonii, C. salina, C. squamata, C. suaveolens, C. subinclusa, C. suksdorfii, C. tuberculata, C. umbellata, C. umbrosa, C. warneri
C. americana, C. approximata, C. azteca, C. boldinghii, C. brachycalyx, C. californica, C. cephalanthi, C. chinensis, C. compacta, C. coryli, C. cuspidata, C. dentatasquamata, C. denticulata, C. draconella, C. epilinum, C. epithymum, C. erosa, C. europaea, C. exaltata, C. glabrior, C. glomerata, C. gronovii, C. harperi, C. howelliana, C. indecora, C. japonica, C. jepsonii, C. legitima, C. leptantha, C. liliputana, C. mitriformis, C. nevadensis, C. obtusiflora, C. occidentalis, C. odontolepis, C. pacifica, C. pentagona, C. plattensis, C. polygonorum, C. rostrata, C. runyonii, C. salina, C. squamata, C. suaveolens, C. subinclusa, C. suksdorfii, C. tuberculata, C. umbellata, C. umbrosa, C. warneri
Synonyms C. arvensis, C. pentagona var. microcalyx, Grammica pentagona C. pentagona var. calycina
Name authority Engelmann: Amer. J. Sci. Arts 43: 340, plate 6, figs. 22–24. (1842) Yuncker: Mem. Torrey Bot. Club 18: 138. (1932)
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