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cryptogramme de Steller, fragile rock-brake, slender rock-brake, slender rock-brake or cliff-brake, Steller's parsley fern, Steller's rock-brake

Alaska parsley fern, Sitka parsley fern, Sitka rockbrake

Stems

creeping, few branched, slender, 1–1.5 mm diam., succulent, brittle;

scales colorless, sparse, transparent-reticulate, ovate, 0.4 × 0.3 mm;

stems shriveling in 2d year following emergence of leaves.

decumbent to erect, much branched from base, stout, 10–20 mm diam. (including hardened, persistent leaf bases);

scales bicolored, dense, broadly lanceolate to linear, to 7 × 2 mm.

Leaves

scattered along stems, ephemeral (dying by late summer), soon shed;

sterile leaves erect, 3–15 cm;

fertile leaves erect, 5–20 cm;

petioles, costae, and costules glabrous.

densely tufted, green over winter, persistent;

fertile leaves erect, 5–25 cm;

sterile leaves spreading, 3–17 cm;

hairs small, appressed, cylindric, scattered along grooves of petioles and along costae and costules of adaxial blade surface.

Petiole

dark brown in proximal 1/2 or less, becoming greenish distally, ca. 1 mm wide when dry, only slightly furrowed, glabrous.

dark brown only on proximal 1/8 or less, green to straw-colored distally, 1–2 mm wide, firm and strawlike, not collapsed;

scales bicolored or ± concolored, becoming sparse distally.

Blade

broadly lanceolate to ovate-lanceolate, all pinnate-pinnatifid to 2-pinnate, herbaceous to membranous, thin;

hydathodes superficial, often poorly developed or absent.

deltate to ovate-lanceolate, somewhat leathery, opaque;

sterile blades dimorphic, 2–3-pinnate or 3–4-pinnate, hydathodes only slightly sunken below leaf surface.

Segments

of sterile leaves ovate-lanceolate to fan-shaped, distal 1/2–1/3 shallowly lobed;

segments of fertile leaves horizontal to ascending, often only partially differentiated from sterile leaves, lanceolate to linear, 8–25 × 2–4 mm;

margins reflexed, forming continuous false indusia.

of less dissected sterile leaves ovate-lanceolate, regularly dentate to incised with 8–16 teeth or lobes;

segments of more finely dissected sterile leaves pinnatifid with 4–8 small, obovate lobes, lobe apices acute;

segments of fertile leaves ascending, strongly differentiated, linear, 3–10 × 1–3 mm;

margins of fertile segments revolute, covering sporangia.

Sporangia

often in discrete sori.

in sori that coalesce at maturity.

2n

= 60.

Cryptogramma stelleri

Cryptogramma sitchensis

Habitat New growth produced in spring, dying by late summer. Sheltered calcareous cliff crevices and rock ledges, typically in coniferous forest or other boreal habitats New growth produced in spring, spores maturing in late summer. sterile leaves green over winter, senescing 2d spring. Cliff crevices and talus slopes, lowland to alpine
Elevation 0–3000 m (0–9800 ft) 0–1800 m (0–5900 ft)
Distribution
from FNA
AK; CO; CT; IA; IL; MA; ME; MI; MN; MT; NH; NJ; NV; NY; OR; PA; UT; VT; WA; WI; WV; WY; AB; BC; NB; NF; NS; NT; ON; PE; QC; YT; Europe in ne former Soviet republics; Asia
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; BC; NT; YT
[BONAP county map]
Discussion

Cryptogramma sitchensis is an allotetraploid species (2n = 120; E. Alverson, unpubl. data) that arose through hybridization between C. acrostichoides and another species, possibly the eastern Asian C. raddeana Fomin. Past difficulties in clearly distinguishing C. sitchensis from C. acrostichoides can be attributed to the frequent occurrence of sterile hybrids where the ranges of the two species overlap.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pteridaceae > Cryptogramma Pteridaceae > Cryptogramma
Sibling taxa
C. acrostichoides, C. cascadensis, C. sitchensis
C. acrostichoides, C. cascadensis, C. stelleri
Synonyms Pteris stelleri Allosorus sitchensis, C. acrostichoides var. sitchensis, C. crispa var. sitchensis
Name authority (S. G. Gmelin) Prantl: in Engler, Bot. Jahrb. Syst. 3: 413. (1882) (Ruprecht) T. Moore: Index Fil. 67. (1857)
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