Cryptogramma cascadensis |
Cryptogramma sitchensis |
|
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Cascade parsely fern, Cascade parsley fern, Cascade rockbrake |
Alaska parsley fern, Sitka parsley fern, Sitka rockbrake |
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Stems | decumbent to erect, much branched from base, stout, 4–8 mm diam. (including hardened, persistent leaf bases); scales often bicolored, dense, broadly lanceolate to linear, to 6 × 2 mm. |
decumbent to erect, much branched from base, stout, 10–20 mm diam. (including hardened, persistent leaf bases); scales bicolored, dense, broadly lanceolate to linear, to 7 × 2 mm. |
Leaves | strongly tufted, deciduous; sterile leaves spreading, 3–20 cm; fertile leaves erect, 5–25 cm; petioles, costae, and costules glabrous. |
densely tufted, green over winter, persistent; fertile leaves erect, 5–25 cm; sterile leaves spreading, 3–17 cm; hairs small, appressed, cylindric, scattered along grooves of petioles and along costae and costules of adaxial blade surface. |
Petiole | green to straw-colored, dark brown only on proximal 1/8 or less, ca. 1 mm wide when dry, collapsing and strongly furrowed; scales bicolored or ± concolored, becoming sparse distally. |
dark brown only on proximal 1/8 or less, green to straw-colored distally, 1–2 mm wide, firm and strawlike, not collapsed; scales bicolored or ± concolored, becoming sparse distally. |
Blade | deltate to ovate-lanceolate, all 2–3-pinnate, herbaceous, thin and translucent when dried, hydathodes superficial. |
deltate to ovate-lanceolate, somewhat leathery, opaque; sterile blades dimorphic, 2–3-pinnate or 3–4-pinnate, hydathodes only slightly sunken below leaf surface. |
Segments | of sterile leaves oblong to fan-shaped, bases cuneate, distal 1/2–1/3 of each segment regularly dentate and often more deeply incised every 2d and 4th tooth; segments of fertile leaves ascending to erect, strongly differentiated from those of sterile leaves, linear, 3–12 × 1–2 mm; fertile segments revolute, covering sporangia. |
of less dissected sterile leaves ovate-lanceolate, regularly dentate to incised with 8–16 teeth or lobes; segments of more finely dissected sterile leaves pinnatifid with 4–8 small, obovate lobes, lobe apices acute; segments of fertile leaves ascending, strongly differentiated, linear, 3–10 × 1–3 mm; margins of fertile segments revolute, covering sporangia. |
Sporangia | in sori that coalesce at maturity. |
in sori that coalesce at maturity. |
2n | = 60. |
|
Cryptogramma cascadensis |
Cryptogramma sitchensis |
|
Habitat | New growth produced in spring, spores maturing in late summer and autumn, leaves dying in autumn. Talus slopes and cliff crevices, often on igneous rocks, typically in relatively mesic subalpine habitats | New growth produced in spring, spores maturing in late summer. sterile leaves green over winter, senescing 2d spring. Cliff crevices and talus slopes, lowland to alpine |
Elevation | 900–3500 m (3000–11500 ft) | 0–1800 m (0–5900 ft) |
Distribution |
CA; ID; MT; OR; WA; BC
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AK; BC; NT; YT |
Discussion | Populations of Cryptogramma cascadensis were previously identified as C. acrostichoides. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cryptogramma sitchensis is an allotetraploid species (2n = 120; E. Alverson, unpubl. data) that arose through hybridization between C. acrostichoides and another species, possibly the eastern Asian C. raddeana Fomin. Past difficulties in clearly distinguishing C. sitchensis from C. acrostichoides can be attributed to the frequent occurrence of sterile hybrids where the ranges of the two species overlap. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pteridaceae > Cryptogramma | Pteridaceae > Cryptogramma |
Sibling taxa | ||
Synonyms | Allosorus sitchensis, C. acrostichoides var. sitchensis, C. crispa var. sitchensis | |
Name authority | E. R. Alverson: Amer. Fern J. 79: 95. (1989) | (Ruprecht) T. Moore: Index Fil. 67. (1857) |
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