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doveweed, goatweed, skunkweed, Texas croton

Alabama croton

Habit Herbs, annual, 2–7(–9) dm, dioecious. Shrubs, 5–35 dm, monoecious.
Stems

loosely branched distally, stellate-hairy.

usually well branched distally, lepidote.

Leaves

not clustered;

stipules absent;

petiole 0.3–2 cm, glands absent at apex;

blade narrowly ovate-oblong to linear-lanceolate, 1–5 × 0.5–2 cm, base truncate to rounded or subcordate, margins entire, apex rounded to acute, abaxial surface pale green, densely whitish appressed stellate-hairy, adaxial surface darker green, less hairy.

clustered at branch tips;

stipules absent;

petiole 0.6–2 cm, glands absent at apex;

blade elliptic, ovate, or oblong, 3–10 × 1.5–5 cm, base rounded to obtuse, margins entire, sometimes ± undulate, apex acute, rounded, or emarginate, abaxial surface silvery or coppery, densely lepidote, adaxial surface green, sparsely lepidote.

Inflorescences

unisexual;

staminate racemes or irregularly branched panicles, 2–8 cm, flowers 10–30;

pistillate racemes, 1–2 cm, flowers 1–6.

bisexual or unisexual, racemes, 2–4.5 cm, staminate flowers 0–15, pistillate flowers 0–10.

Pedicels

staminate 2–3 mm, pistillate 1–3 mm.

staminate 2.2–4 mm, pistillate 2.2–7.5 mm (7–11 mm in fruit).

Staminate flowers

sepals 5, 1–2 mm, abaxial surface densely whitish appressed stellate-hairy;

petals 0;

stamens 8–12.

sepals 5, 1.1–2.9 mm, abaxial surface lepidote;

petals 5, oblong-ovate, 2–3.1 mm, abaxial surface glabrous except margins stellate-ciliate;

stamens 10–22.

Pistillate flowers

sepals 5, equal, 1–1.5 mm, margins entire, apex incurved, abaxial surface densely stipitate-stellate-hairy;

petals 0;

ovary 3-locular;

styles 3, 1–2 mm, multifid, terminal segments 12–32+.

sepals 5, equal, 2–4.5 mm, margins entire, apex incurved, abaxial surface lepidote;

petals 5, pale green, ovate, 2–3.5 mm;

ovary 3-locular;

styles 3, 2–5 mm, usually unbranched, rarely 2-fid, terminal segments 3 (or 6).

Capsules

5–8 × 4–5.5 mm, verrucose;

columella 3-winged.

1.6–2.5 × 2–3 mm, smooth;

columella 3-angled.

Seeds

3.5–4 × 2.5–3 mm, shiny.

6.7–8 × 5.2–6 mm, shiny.

2n

= 28.

Croton texensis

Croton alabamensis

Phenology Flowering Jun–Nov.
Habitat Prairies, sandy creek beds, old fields, canyons, disturbed areas.
Elevation 50–2000 m. (200–6600 ft.)
Distribution
from FNA
AL; AZ; CO; DE; FL; IA; IL; KS; MD; MO; NE; NM; OK; SD; TX; UT; WI; WV; WY; Mexico (Chihuahua, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Croton texensis, despite being annual, grows larger than the related perennial C. dioicus. Croton texensis has verrucose fruits similar to those of the closely related C. parksii.

There is a single specimen of Croton texensis from Massachusetts, collected at a dump in Boston in 1890, but the species did not become established there.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

B. W. van Ee et al. (2006) examined populations of Croton alabamensis using DNA sequence and AFLP data, and their results supported the recognition of two varieties and an isolated position in the genus. Buds develop in the summer and fall for the following spring's flowers.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blade abaxial surfaces silvery, scales mostly unpigmented or light amber, some dark blackish brown scales sometimes present; inflorescences 9–18-flowered, producing 0–11 fruits; Alabama.
var. alabamensis
1. Leaf blade abaxial surfaces coppery, some scales unpigmented, others with dark reddish brown center and reddish amber rays; inflorescences 6–14-flowered, producing 0–6 fruits; Texas.
var. texensis
Source FNA vol. 12, p. 224. FNA vol. 12, p. 210.
Parent taxa Euphorbiaceae > Croton Euphorbiaceae > Croton
Sibling taxa
C. alabamensis, C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. glandulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. wigginsii
C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. glandulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
Subordinate taxa
C. alabamensis var. alabamensis, C. alabamensis var. texensis
Synonyms Hendecandra texensis, C. luteovirens, C. texensis var. utahensis, C. virens
Name authority (Klotzsch) Müller Arg.: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 15(2): 692. (1866) E. A. Smith ex Chapman: Fl. South. U.S. ed. 2, 648. (1883)
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