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beach tea, gulf croton, hierba de jabalí

sand croton, sand or tooth-leaf or Tropic croton, tooth-leaf croton, Tropic croton, vente conmigo

Habit Shrubs, 3–10 dm, monoecious or sometimes appearing dioecious. Herbs, annual, 1–12 dm, monoecious.
Stems

trichotomously branching, stellate-hairy to stellate-lepidote.

much branched distally, usually coarsely stellate-hairy, rarely glabrescent.

Leaves

not clustered;

stipules absent;

petiole 1–4 cm, 1/2 to equal blade length, glands absent at apex;

blade broadly elliptic to suborbiculate, 2–5 × 1.5–4 cm, much less than 2 times as long as wide, base rounded to truncate, margins entire, apex obtuse to rounded, abaxial surface pale grayish green, stellate-lepidote, adaxial surface slightly darker green, stellate-lepidote.

not clustered;

stipules linear-subulate, glandular or not, to 0.5 mm or absent;

petiole 0.2–1 cm, glands at apex 2, yellow, sessile to shortly stipitate, cuplike;

blade ovate proximally, oblong-lanceolate distally, 0.6–3.5(–7) × 0.3–1.5(–3) cm, base obtuse to truncate, margins coarsely crenate to serrate-dentate, apex obtuse to rounded, both surfaces green, stellate-hairy, glabrate, or rarely glabrous.

Inflorescences

unisexual or bisexual, racemes, 1–4 cm, staminate flowers 3–7, pistillate flowers 1–3.

bisexual, racemes, 1–3 cm, staminate flowers 10–20, pistillate flowers 1–4.

Pedicels

staminate 2–4 mm, pistillate 0–1 mm.

staminate 0.8–2 mm, pistillate 0–5 mm.

Staminate flowers

sepals 5–6, 2.5 mm, abaxial surface stellate-lepidote;

petals 0;

stamens 10–13.

sepals 5, 0.8–1.2 mm, abaxial surface stellate-hairy;

petals 5, oblanceolate, 1–1.3 mm, abaxial surface glabrous except margins ciliate;

stamens 7–13.

Pistillate flowers

sepals 5, equal, 3–3.5 mm, margins entire, apex incurved, abaxial surface stellate-lepidote;

petals 0;

ovary 3-locular;

styles 3, 1–2 mm, multifid, terminal segments 12–24.

sepals 5, subequal, 6–7.5 mm, margins entire, apex straight to slightly incurved, abaxial surface glabrous except stellate-hairy apically;

petals 0 or 5, rudimentary;

ovary 3-locular;

styles 3, 1–2.5 mm, deeply 2-fid, terminal segments 6.

Capsules

5–8 × 7–9 mm, smooth;

columella 3-winged.

3.5–6 × 4–5 mm, smooth;

columella 3-angled.

Seeds

4.5–6 × 3.7–4.5 mm, dull.

3–4 × 2–2.5 mm, shiny.

2n

= 28.

= 16.

Croton punctatus

Croton glandulosus

Phenology Flowering year-round.
Habitat Beaches, dunes.
Elevation 0–20 m. (0–100 ft.)
Distribution
from FNA
AL; FL; GA; LA; MS; NC; SC; TX; e Mexico; Central America; West Indies; n South America
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; DE; FL; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; NC; NE; NJ; NM; OH; OK; PA; SC; TN; TX; VA; WI; WV; Mexico; Central America; South America; West Indies
[WildflowerSearch map]
[BONAP county map]
Discussion

A report of Croton punctatus from Pennsylvania apparently was based on a transient appearance on ballast (E. T. Wherry et al. 1979).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties ca. 20 (5 in the flora).

Croton glandulosus is widespread in the New World, with a complex pattern of variation. The classification here follows B. W. van Ee et al. (2009).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades glabrate or sparsely to moderately stellate-hairy.
→ 2
2. Plants 2–12 dm; leaf blades 2–7 cm.
var. septentrionalis
2. Plants 1–2 dm; leaf blades 0.6–2(–3) cm.
→ 3
3. Leaf blades glabrate, bases markedly 3-veined; petiole apical glands sessile; Florida.
var. floridanus
3. Leaf blades sparsely stellate-hairy, bases obscurely 3-veined; petiole apical glands stipitate; Kansas, New Mexico, Oklahoma, Texas.
var. lindheimeri
1. Leaf blades moderately to densely stellate-hairy.
→ 4
4. Leaf blades firm-thick, length mostly 2 times width or less, marginal teeth rounded; petiole apical glands sessile.
var. arenicola
4. Leaf blades membranous, length mostly more than 2 times width, marginal teeth pointed; petiole apical glands sessile or stipitate.
→ 5
5. Leaf blades 1–2(–3) × 0.3–0.8(–1.3) cm; petiole apical glands stipitate, circular when dry, 0.1–0.4 mm diam.
var. lindheimeri
5. Leaf blades 2–7 × 0.7–3 cm; petiole apical glands sessile, wavy-wrinkled when dry, 0.5–0.8 mm diam.
→ 6
6. Stems densely stellate-hairy, hairs spreading, radii unequal; leaf blades densely stellate-villous.
var. pubentissimus
6. Stems moderately stellate-hairy, hairs appressed, radii equal; leaf blades moderately stellate-hairy.
var. septentrionalis
Source FNA vol. 12, p. 222. FNA vol. 12, p. 215.
Parent taxa Euphorbiaceae > Croton Euphorbiaceae > Croton
Sibling taxa
C. alabamensis, C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. glandulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
C. alabamensis, C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
Subordinate taxa
C. glandulosus var. arenicola, C. glandulosus var. floridanus, C. glandulosus var. lindheimeri, C. glandulosus var. pubentissimus, C. glandulosus var. septentrionalis
Synonyms C. disjunctiflorus, C. maritimus, C. plukenetii Decarinium glandulosum, Geiseleria glandulosa, Oxydectes glandulosa
Name authority Jacquin: Collectanea 1: 166. (1787) — (as puntatum) Linnaeus: Syst. Nat. ed. 10, 2: 1275. (1759) — (as glandulosum)
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