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woolly croton

Cortez's croton, palillo

Habit Herbs, annual, 5–15 dm, monoecious; stems, leaves, and buds whitish-hairy when young, becoming glabrate. Shrubs, 10–30 dm, dioecious.
Stems

well branched distally, stellate-hairy.

di- or trichotomously branched, stellate-hairy, young stems often blackened by fungus.

Leaves

not clustered;

stipules linear, 2–7 mm;

petiole 0.5–5 cm, glands absent at apex;

blade ovate-lanceolate, 3–10 × 1–5 cm, base cordate to rounded, margins entire, apex acute, abaxial surface pale green, not appearing brown-dotted, no stellate hairs with brown centers, adaxial surface darker green, both stellate-hairy.

not clustered;

stipules rudimentary;

petiole 1–1.5 cm, 1/6–1/3 leaf blade length, glands absent at apex;

blade narrowly obovate, oblong, elliptic, or lanceolate, 3–10 × 1–5 cm, mostly less than 3.5 times as long as wide, base subacute to rounded, margins minutely denticulate, apex acute or acuminate, abaxial surface light green, densely ochraceous stellate-hispid when young, pale stellate-hairy when mature, adaxial surface darker green, glabrous.

Inflorescences

bisexual, racemes, 2–4 cm, staminate flowers 3–10, pistillate flowers 4–8.

unisexual, racemes;

staminate 7–20 cm, flowers 10–30;

pistillate 1–5 cm, flowers 8–20(–30).

Pedicels

staminate 2–4 mm, pistillate 1–2 mm.

staminate 0–1 mm, pistillate 0–0.5 mm.

Staminate flowers

sepals (4–)5, 1–2 mm, abaxial surface stellate-hairy;

petals 5, linear-oblong-lanceolate, 1–1.5 mm, abaxial surface stellate-hairy;

stamens 9–12.

sepals 5, 2.5 mm, abaxial surface stellate-hairy;

petals 5, oblong, 3 mm, abaxial surface glabrous except margins ciliate basally;

stamens 12–16.

Pistillate flowers

sepals 7–8, subequal, 3–6 mm, margins entire, apex straight to slightly incurved, abaxial surface whitish appressed-tomentose;

petals 0;

ovary 3-locular;

styles 3, 3–4 mm, 4-fid, terminal segments 12.

sepals 5, equal, 1–1.5 mm, margins entire, apex straight to slightly incurved, abaxial surface stellate-hispid;

petals rudimentary;

ovary 3-locular;

styles 3, 4–5 mm, 2-fid, terminal segments 6.

Capsules

6–8 × 6–7 mm, smooth;

columella tipped with 3-pronged grappling hooklike appendage.

5–6 × 4.5–5.5 mm, smooth;

columella apex with 3 rounded, inflated lobes.

Seeds

4–5 × 2–2.5 mm, shiny.

4–5 × 3–3.5 mm, shiny (with silvery sheen).

2n

= 20.

Croton heptalon

Croton cortesianus

Phenology Flowering May–Dec. Flowering Feb–Oct, possibly year-round.
Habitat Beaches, coastal dunes, roadsides. Scrub forests.
Elevation 0–50 m. (0–200 ft.) 0–50 m. (0–200 ft.)
Distribution
from FNA
TX; e Mexico
[BONAP county map]
from FNA
TX; e Mexico; se Mexico; Central America
[BONAP county map]
Discussion

Morphological differences among Croton heptalon and its multifid-styled relatives in sect. Heptallon, especially C. capitatus and C. lindheimeri, can be quite subtle. In general, C. heptalon can be distinguished from C. capitatus by its more elongate pistillate part of the inflorescence, non-recurving sepal tips in the pistillate flowers, and more cordate leaf bases on larger basal leaves. Whitish pubescence on its young growth and styles that branch once into four terminal segments distinguish C. heptalon from C. lindheimeri.

Croton muelleri J. M. Coulter, which is an illegitimate name, pertains here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Croton cortesianus is known in the flora area only from Cameron, Hidalgo, and Starr counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 217. FNA vol. 12, p. 213.
Parent taxa Euphorbiaceae > Croton Euphorbiaceae > Croton
Sibling taxa
C. alabamensis, C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. glandulosus, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
C. alabamensis, C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. glandulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
Synonyms Oxydectes heptalon, C. albinoides, C. capitatus var. albinoides, C. engelmannii var. albinoides, C. muelleri var. albinoides, Heptallon aromaticum C. chichenensis, C. trichocarpus
Name authority (Kuntze) B. W. van Ee & P. E. Berry: Syst. Bot. 35: 159. (2010) Kunth: in A. von Humboldt et al., Nov. Gen. Sp. 2(fol.): 66; 2(qto.): 83. (1817)
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