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sand croton, sand or tooth-leaf or Tropic croton, tooth-leaf croton, Tropic croton, vente conmigo

scented croton

Habit Herbs, annual, 1–12 dm, monoecious. Shrubs, 2–5 dm, monoecious.
Stems

much branched distally, usually coarsely stellate-hairy, rarely glabrescent.

dichotomously much branched proximally, coarsely stellate-tomentose.

Leaves

not clustered;

stipules linear-subulate, glandular or not, to 0.5 mm or absent;

petiole 0.2–1 cm, glands at apex 2, yellow, sessile to shortly stipitate, cuplike;

blade ovate proximally, oblong-lanceolate distally, 0.6–3.5(–7) × 0.3–1.5(–3) cm, base obtuse to truncate, margins coarsely crenate to serrate-dentate, apex obtuse to rounded, both surfaces green, stellate-hairy, glabrate, or rarely glabrous.

often clustered near inflorescences;

stipules each 5–10 glandular papillae, 0.2–0.5 mm;

petiole 0.5–1.5 cm, 1/4 leaf blade length, glands absent at apex;

blade obovate or ovate to broadly elliptic, 2–5.4 × 1–3.6 cm, base obtuse, margins entire, apex obtuse to acute, abaxial surface pale green to cream, adaxial surface darker green, both densely stellate-hairy.

Inflorescences

bisexual, racemes, 1–3 cm, staminate flowers 10–20, pistillate flowers 1–4.

bisexual or staminate, racemes, 1–3 cm, staminate flowers 6–10, pistillate flowers 1–3.

Pedicels

staminate 0.8–2 mm, pistillate 0–5 mm.

staminate 3–5 mm, pistillate 1–2 mm.

Staminate flowers

sepals 5, 0.8–1.2 mm, abaxial surface stellate-hairy;

petals 5, oblanceolate, 1–1.3 mm, abaxial surface glabrous except margins ciliate;

stamens 7–13.

sepals 5, 2 mm, abaxial surface stellate-hairy;

petals 5, spatulate or obovate, 0.4–0.6 mm, abaxial surface densely villous at base and along margins, otherwise glabrous;

stamens 12–16.

Pistillate flowers

sepals 5, subequal, 6–7.5 mm, margins entire, apex straight to slightly incurved, abaxial surface glabrous except stellate-hairy apically;

petals 0 or 5, rudimentary;

ovary 3-locular;

styles 3, 1–2.5 mm, deeply 2-fid, terminal segments 6.

sepals 5, equal, 2.5 mm, margins entire, apex straight to slightly incurved, abaxial surface stellate-hairy;

petals glandular papillae;

ovary 3-locular;

styles 3, 4–6 mm, 2-fid to base, terminal segments 6.

Capsules

3.5–6 × 4–5 mm, smooth;

columella 3-angled.

7–10 × 6–8 mm, smooth;

columella apex with 3 rounded, inflated lobes.

Seeds

3–4 × 2–2.5 mm, shiny.

5.5–7 × 4–5 mm, shiny.

2n

= 16.

Croton glandulosus

Croton suaveolens

Phenology Flowering Apr–Nov.
Habitat Rocky slopes, foothills.
Elevation 1500–2000 m. (4900–6600 ft.)
Distribution
from FNA
AL; AR; DE; FL; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; NC; NE; NJ; NM; OH; OK; PA; SC; TN; TX; VA; WI; WV; Mexico; Central America; South America; West Indies
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; Mexico (Coahuila, Chihuahua, Nuevo León)
[BONAP county map]
Discussion

Varieties ca. 20 (5 in the flora).

Croton glandulosus is widespread in the New World, with a complex pattern of variation. The classification here follows B. W. van Ee et al. (2009).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Croton suaveolens is found in the flora area only in trans-Pecos Texas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades glabrate or sparsely to moderately stellate-hairy.
→ 2
2. Plants 2–12 dm; leaf blades 2–7 cm.
var. septentrionalis
2. Plants 1–2 dm; leaf blades 0.6–2(–3) cm.
→ 3
3. Leaf blades glabrate, bases markedly 3-veined; petiole apical glands sessile; Florida.
var. floridanus
3. Leaf blades sparsely stellate-hairy, bases obscurely 3-veined; petiole apical glands stipitate; Kansas, New Mexico, Oklahoma, Texas.
var. lindheimeri
1. Leaf blades moderately to densely stellate-hairy.
→ 4
4. Leaf blades firm-thick, length mostly 2 times width or less, marginal teeth rounded; petiole apical glands sessile.
var. arenicola
4. Leaf blades membranous, length mostly more than 2 times width, marginal teeth pointed; petiole apical glands sessile or stipitate.
→ 5
5. Leaf blades 1–2(–3) × 0.3–0.8(–1.3) cm; petiole apical glands stipitate, circular when dry, 0.1–0.4 mm diam.
var. lindheimeri
5. Leaf blades 2–7 × 0.7–3 cm; petiole apical glands sessile, wavy-wrinkled when dry, 0.5–0.8 mm diam.
→ 6
6. Stems densely stellate-hairy, hairs spreading, radii unequal; leaf blades densely stellate-villous.
var. pubentissimus
6. Stems moderately stellate-hairy, hairs appressed, radii equal; leaf blades moderately stellate-hairy.
var. septentrionalis
Source FNA vol. 12, p. 215. FNA vol. 12, p. 224.
Parent taxa Euphorbiaceae > Croton Euphorbiaceae > Croton
Sibling taxa
C. alabamensis, C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
C. alabamensis, C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. glandulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. texensis, C. wigginsii
Subordinate taxa
C. glandulosus var. arenicola, C. glandulosus var. floridanus, C. glandulosus var. lindheimeri, C. glandulosus var. pubentissimus, C. glandulosus var. septentrionalis
Synonyms Decarinium glandulosum, Geiseleria glandulosa, Oxydectes glandulosa
Name authority Linnaeus: Syst. Nat. ed. 10, 2: 1275. (1759) — (as glandulosum) Torrey: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 194. (1859)
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