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sand croton, sand or tooth-leaf or Tropic croton, tooth-leaf croton, Tropic croton, vente conmigo

one-seed croton, prairie tea

Habit Herbs, annual, 1–12 dm, monoecious. Herbs, annual, 2–5 dm, monoecious.
Stems

much branched distally, usually coarsely stellate-hairy, rarely glabrescent.

dichotomously branched from near base, stellate-hairy, some stellate hairs with dark brown centers.

Leaves

not clustered;

stipules linear-subulate, glandular or not, to 0.5 mm or absent;

petiole 0.2–1 cm, glands at apex 2, yellow, sessile to shortly stipitate, cuplike;

blade ovate proximally, oblong-lanceolate distally, 0.6–3.5(–7) × 0.3–1.5(–3) cm, base obtuse to truncate, margins coarsely crenate to serrate-dentate, apex obtuse to rounded, both surfaces green, stellate-hairy, glabrate, or rarely glabrous.

sometimes clustered near inflorescences;

stipules glandlike, 0.1–0.3 mm;

petiole 0.3–1.5 cm, glands absent at apex;

blade ovate-oblong to nearly round (proximal) to narrowly elliptic (distal), 1–3.5 × 0.5–3 cm, base obtuse to rounded or truncate, margins entire, apex rounded to acute, abaxial surface pale green, sparsely whitish stellate-hairy and appearing brown-dotted, some hairs with dark brown centers, adaxial surface darker green, densely stellate-hairy, hairs without brown centers.

Inflorescences

bisexual, racemes, 1–3 cm, staminate flowers 10–20, pistillate flowers 1–4.

bisexual or sometimes pistillate, congested racemes, 0.3–1 cm, staminate flowers 3–10, pistillate flowers 1–2(–5).

Pedicels

staminate 0.8–2 mm, pistillate 0–5 mm.

staminate 0.5–2 mm, pistillate 1–2.5 mm (2–3 mm and recurved in fruit).

Staminate flowers

sepals 5, 0.8–1.2 mm, abaxial surface stellate-hairy;

petals 5, oblanceolate, 1–1.3 mm, abaxial surface glabrous except margins ciliate;

stamens 7–13.

sepals 3–5, 0.7–1 mm, abaxial surface stellate-hairy;

petals 3–5, narrowly elliptic-oblanceolate, 0.7–1 mm, abaxial surface glabrous except margins villous;

stamens 3–5.

Pistillate flowers

sepals 5, subequal, 6–7.5 mm, margins entire, apex straight to slightly incurved, abaxial surface glabrous except stellate-hairy apically;

petals 0 or 5, rudimentary;

ovary 3-locular;

styles 3, 1–2.5 mm, deeply 2-fid, terminal segments 6.

sepals 5, subequal, 1.5–2 mm, margins entire, apex straight to slightly incurved, abaxial surface stellate-hairy, some hairs with dark brown centers;

petals 0;

ovary 2-locular, 1 fertile;

styles 2, 0.8–1.2(–1.5) mm, 2-fid to base, terminal segments 4.

Capsules

3.5–6 × 4–5 mm, smooth;

columella 3-angled.

appearing follicular (1-seeded), 3.5–4.5 × 1.8–2.2 mm, smooth;

columella ± curved, apparently deciduous upon dehiscence of capsule.

Seeds

3–4 × 2–2.5 mm, shiny.

2.5–3.3 × 2–2.5 mm, shiny.

2n

= 16.

= 20.

Croton glandulosus

Croton monanthogynus

Phenology Flowering May–Nov.
Habitat Prairies, sandstone and limestone glades, thinly wooded bluffs, fallow fields, other disturbed habitats.
Elevation 0–1000 m. (0–3300 ft.)
Distribution
from FNA
AL; AR; DE; FL; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; NC; NE; NJ; NM; OH; OK; PA; SC; TN; TX; VA; WI; WV; Mexico; Central America; South America; West Indies
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; GA; IA; IL; IN; KS; KY; LA; MD; MI; MO; MS; NC; NE; NM; OH; OK; PA; SC; TN; TX; VA; WI; WV; Mexico (Coahuila, Nuevo León, San Luis Potosí, Tamaulipas)
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties ca. 20 (5 in the flora).

Croton glandulosus is widespread in the New World, with a complex pattern of variation. The classification here follows B. W. van Ee et al. (2009).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Croton monanthogynus may be adventive in the northernmost states.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades glabrate or sparsely to moderately stellate-hairy.
→ 2
2. Plants 2–12 dm; leaf blades 2–7 cm.
var. septentrionalis
2. Plants 1–2 dm; leaf blades 0.6–2(–3) cm.
→ 3
3. Leaf blades glabrate, bases markedly 3-veined; petiole apical glands sessile; Florida.
var. floridanus
3. Leaf blades sparsely stellate-hairy, bases obscurely 3-veined; petiole apical glands stipitate; Kansas, New Mexico, Oklahoma, Texas.
var. lindheimeri
1. Leaf blades moderately to densely stellate-hairy.
→ 4
4. Leaf blades firm-thick, length mostly 2 times width or less, marginal teeth rounded; petiole apical glands sessile.
var. arenicola
4. Leaf blades membranous, length mostly more than 2 times width, marginal teeth pointed; petiole apical glands sessile or stipitate.
→ 5
5. Leaf blades 1–2(–3) × 0.3–0.8(–1.3) cm; petiole apical glands stipitate, circular when dry, 0.1–0.4 mm diam.
var. lindheimeri
5. Leaf blades 2–7 × 0.7–3 cm; petiole apical glands sessile, wavy-wrinkled when dry, 0.5–0.8 mm diam.
→ 6
6. Stems densely stellate-hairy, hairs spreading, radii unequal; leaf blades densely stellate-villous.
var. pubentissimus
6. Stems moderately stellate-hairy, hairs appressed, radii equal; leaf blades moderately stellate-hairy.
var. septentrionalis
Source FNA vol. 12, p. 215. FNA vol. 12, p. 220.
Parent taxa Euphorbiaceae > Croton Euphorbiaceae > Croton
Sibling taxa
C. alabamensis, C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
C. alabamensis, C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. glandulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
Subordinate taxa
C. glandulosus var. arenicola, C. glandulosus var. floridanus, C. glandulosus var. lindheimeri, C. glandulosus var. pubentissimus, C. glandulosus var. septentrionalis
Synonyms Decarinium glandulosum, Geiseleria glandulosa, Oxydectes glandulosa Engelmannia nuttalliana, Gynamblosis monanthogyna, Heptallon ellipticum, Oxydectes monanthogyna
Name authority Linnaeus: Syst. Nat. ed. 10, 2: 1275. (1759) — (as glandulosum) Michaux: Fl. Bor.-Amer. 2: 215. (1803) — (as monanthogynum)
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