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sand croton, sand or tooth-leaf or Tropic croton, tooth-leaf croton, Tropic croton, vente conmigo

Michaux's croton, narrowleaf rushfoil, rushfoil

Habit Herbs, annual, 1–12 dm, monoecious. Herbs, annual, 1–9 dm, monoecious.
Stems

much branched distally, usually coarsely stellate-hairy, rarely glabrescent.

usually single from base, much branched from near middle, densely stellate-lepidote appearing silvery.

Leaves

not clustered;

stipules linear-subulate, glandular or not, to 0.5 mm or absent;

petiole 0.2–1 cm, glands at apex 2, yellow, sessile to shortly stipitate, cuplike;

blade ovate proximally, oblong-lanceolate distally, 0.6–3.5(–7) × 0.3–1.5(–3) cm, base obtuse to truncate, margins coarsely crenate to serrate-dentate, apex obtuse to rounded, both surfaces green, stellate-hairy, glabrate, or rarely glabrous.

not clustered;

stipules rudimentary;

petiole 0.1–0.5 cm, glands absent at apex;

blade lanceolate-linear to ovate-lanceolate, 1.5–4 × 0.1–0.4(–1.5) cm, base acute to cuneate, margins entire, apex acute, abaxial surface silvery, stellate-lepidote, adaxial surface green, sparsely stellate-hairy, hairs with few radii.

Inflorescences

bisexual, racemes, 1–3 cm, staminate flowers 10–20, pistillate flowers 1–4.

bisexual, spikes, 0.3–3 cm, staminate flowers 1–12, pistillate flowers 1–6.

Pedicels

staminate 0.8–2 mm, pistillate 0–5 mm.

pistillate absent, staminate 0–1 mm.

Staminate flowers

sepals 5, 0.8–1.2 mm, abaxial surface stellate-hairy;

petals 5, oblanceolate, 1–1.3 mm, abaxial surface glabrous except margins ciliate;

stamens 7–13.

sepals 5, 1 mm, abaxial surface lepidote;

petals 5, spatulate, 0.6–1 mm, abaxial surface glabrous except margins villous;

stamens 5–7.

Pistillate flowers

sepals 5, subequal, 6–7.5 mm, margins entire, apex straight to slightly incurved, abaxial surface glabrous except stellate-hairy apically;

petals 0 or 5, rudimentary;

ovary 3-locular;

styles 3, 1–2.5 mm, deeply 2-fid, terminal segments 6.

sepals (4–)5, equal, 1–1.5 mm, margins entire, apex straight, abaxial surface lepidote;

petals 0;

ovary 1-locular;

styles 3, 0.5–0.8 mm, minutely 2-fid, terminal segments 3 or 6.

Capsules

3.5–6 × 4–5 mm, smooth;

columella 3-angled.

Achenes

2.5–3 × 1–1.2 mm, smooth;

columella absent.

Seeds

3–4 × 2–2.5 mm, shiny.

2–2.5 × 1 mm, surface not visible because seed coat adherent to fruit wall.

2n

= 16.

Croton glandulosus

Croton michauxii

Distribution
from FNA
AL; AR; DE; FL; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; NC; NE; NJ; NM; OH; OK; PA; SC; TN; TX; VA; WI; WV; Mexico; Central America; South America; West Indies
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MD; MO; MS; NC; NJ; OK; PA; SC; TN; TX; VA
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties ca. 20 (5 in the flora).

Croton glandulosus is widespread in the New World, with a complex pattern of variation. The classification here follows B. W. van Ee et al. (2009).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades glabrate or sparsely to moderately stellate-hairy.
→ 2
2. Plants 2–12 dm; leaf blades 2–7 cm.
var. septentrionalis
2. Plants 1–2 dm; leaf blades 0.6–2(–3) cm.
→ 3
3. Leaf blades glabrate, bases markedly 3-veined; petiole apical glands sessile; Florida.
var. floridanus
3. Leaf blades sparsely stellate-hairy, bases obscurely 3-veined; petiole apical glands stipitate; Kansas, New Mexico, Oklahoma, Texas.
var. lindheimeri
1. Leaf blades moderately to densely stellate-hairy.
→ 4
4. Leaf blades firm-thick, length mostly 2 times width or less, marginal teeth rounded; petiole apical glands sessile.
var. arenicola
4. Leaf blades membranous, length mostly more than 2 times width, marginal teeth pointed; petiole apical glands sessile or stipitate.
→ 5
5. Leaf blades 1–2(–3) × 0.3–0.8(–1.3) cm; petiole apical glands stipitate, circular when dry, 0.1–0.4 mm diam.
var. lindheimeri
5. Leaf blades 2–7 × 0.7–3 cm; petiole apical glands sessile, wavy-wrinkled when dry, 0.5–0.8 mm diam.
→ 6
6. Stems densely stellate-hairy, hairs spreading, radii unequal; leaf blades densely stellate-villous.
var. pubentissimus
6. Stems moderately stellate-hairy, hairs appressed, radii equal; leaf blades moderately stellate-hairy.
var. septentrionalis
1. Inflorescences 1–3 cm; achenes usually 3–6, densely stellate-lepidote, hairs ± spreading, radii separate most of length, umbos usually sharp, sometimes rounded; leaf blade adaxial surface hairs not overlapping.
var. michauxii
1. Inflorescences usually less than 1 cm; achenes 1–2, sparsely stellate-lepidote, hairs appressed, radii joined most or all of length, umbos usually blunt, rarely sharp; leaf blade adaxial surface hairs overlapping near midvein.
var. elliptica
Source FNA vol. 12, p. 215. FNA vol. 12, p. 220.
Parent taxa Euphorbiaceae > Croton Euphorbiaceae > Croton
Sibling taxa
C. alabamensis, C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
C. alabamensis, C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. glandulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
Subordinate taxa
C. glandulosus var. arenicola, C. glandulosus var. floridanus, C. glandulosus var. lindheimeri, C. glandulosus var. pubentissimus, C. glandulosus var. septentrionalis
C. michauxii var. elliptica, C. michauxii var. michauxii
Synonyms Decarinium glandulosum, Geiseleria glandulosa, Oxydectes glandulosa Crotonopsis linearis
Name authority Linnaeus: Syst. Nat. ed. 10, 2: 1275. (1759) — (as glandulosum) G. L. Webster: Novon 2: 270. (1992)
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