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sand croton, sand or tooth-leaf or Tropic croton, tooth-leaf croton, Tropic croton, vente conmigo

grannybush, pineland croton

Habit Herbs, annual, 1–12 dm, monoecious. Shrubs, 10–20 dm, dioecious.
Stems

much branched distally, usually coarsely stellate-hairy, rarely glabrescent.

well branched from base, stellate-hairy.

Leaves

not clustered;

stipules linear-subulate, glandular or not, to 0.5 mm or absent;

petiole 0.2–1 cm, glands at apex 2, yellow, sessile to shortly stipitate, cuplike;

blade ovate proximally, oblong-lanceolate distally, 0.6–3.5(–7) × 0.3–1.5(–3) cm, base obtuse to truncate, margins coarsely crenate to serrate-dentate, apex obtuse to rounded, both surfaces green, stellate-hairy, glabrate, or rarely glabrous.

not clustered;

stipules rudimentary;

petiole 0.3–1 cm, 1/10–1/7 leaf blade length, often with 2 sessile or stipitate glands at apex;

blade linear to narrowly oblong, 3–7 × 0.3–1.5 cm, mostly more than 4 times as long as wide, base cuneate to obtuse, margins entire, apex obtuse to acute, abaxial surface whitish yellow, densely appressed stellate-hairy, adaxial surface green, glabrous or minutely stellate-puberulent.

Inflorescences

bisexual, racemes, 1–3 cm, staminate flowers 10–20, pistillate flowers 1–4.

unisexual, racemes;

staminate 4–10 cm, flowers 10–30;

pistillate 3–5 cm, flowers 5–12.

Pedicels

staminate 0.8–2 mm, pistillate 0–5 mm.

staminate 1–2 mm, pistillate 2–3 mm.

Staminate flowers

sepals 5, 0.8–1.2 mm, abaxial surface stellate-hairy;

petals 5, oblanceolate, 1–1.3 mm, abaxial surface glabrous except margins ciliate;

stamens 7–13.

sepals 5(–6), 2–2.5 mm, abaxial surface stellate-hairy;

petals 5–6, spatulate, obtuse, 1.5–2.5 mm, abaxial surface glabrous except margins ciliate;

stamens 12–17.

Pistillate flowers

sepals 5, subequal, 6–7.5 mm, margins entire, apex straight to slightly incurved, abaxial surface glabrous except stellate-hairy apically;

petals 0 or 5, rudimentary;

ovary 3-locular;

styles 3, 1–2.5 mm, deeply 2-fid, terminal segments 6.

sepals 5, equal, 2.5–3.5 mm, margins entire, apex incurved, abaxial surface canescent;

petals rudimentary or 0;

ovary 3-locular;

styles 3, 2–3 mm, 2-fid, terminal segments 6.

Capsules

3.5–6 × 4–5 mm, smooth;

columella 3-angled.

5–7 × 4–5 mm, smooth;

columella with 3 rounded, inflated lobes.

Seeds

3–4 × 2–2.5 mm, shiny.

3–4 × 1.5–2 mm, dull.

2n

= 16.

Croton glandulosus

Croton linearis

Phenology Flowering year-round.
Habitat Rocky limestone pinelands, coastal areas.
Elevation 0–30 m. (0–100 ft.)
Distribution
from FNA
AL; AR; DE; FL; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; NC; NE; NJ; NM; OH; OK; PA; SC; TN; TX; VA; WI; WV; Mexico; Central America; South America; West Indies
[WildflowerSearch map]
[BONAP county map]
from FNA
FL; West Indies
[WildflowerSearch map]
Discussion

Varieties ca. 20 (5 in the flora).

Croton glandulosus is widespread in the New World, with a complex pattern of variation. The classification here follows B. W. van Ee et al. (2009).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Croton linearis in the flora area is found only in Martin, Miami-Dade, Monroe (Keys only), Palm Beach, and St. Lucie counties.

There has been some confusion in the past with the application of the name C. cascarilla (Linnaeus) Linnaeus to this taxon, but C. cascarilla is now considered to be a synonym of the West Indian C. eluteria (Linnaeus) W. Wright (B. W. van Ee and P. E. Berry 2010b).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades glabrate or sparsely to moderately stellate-hairy.
→ 2
2. Plants 2–12 dm; leaf blades 2–7 cm.
var. septentrionalis
2. Plants 1–2 dm; leaf blades 0.6–2(–3) cm.
→ 3
3. Leaf blades glabrate, bases markedly 3-veined; petiole apical glands sessile; Florida.
var. floridanus
3. Leaf blades sparsely stellate-hairy, bases obscurely 3-veined; petiole apical glands stipitate; Kansas, New Mexico, Oklahoma, Texas.
var. lindheimeri
1. Leaf blades moderately to densely stellate-hairy.
→ 4
4. Leaf blades firm-thick, length mostly 2 times width or less, marginal teeth rounded; petiole apical glands sessile.
var. arenicola
4. Leaf blades membranous, length mostly more than 2 times width, marginal teeth pointed; petiole apical glands sessile or stipitate.
→ 5
5. Leaf blades 1–2(–3) × 0.3–0.8(–1.3) cm; petiole apical glands stipitate, circular when dry, 0.1–0.4 mm diam.
var. lindheimeri
5. Leaf blades 2–7 × 0.7–3 cm; petiole apical glands sessile, wavy-wrinkled when dry, 0.5–0.8 mm diam.
→ 6
6. Stems densely stellate-hairy, hairs spreading, radii unequal; leaf blades densely stellate-villous.
var. pubentissimus
6. Stems moderately stellate-hairy, hairs appressed, radii equal; leaf blades moderately stellate-hairy.
var. septentrionalis
Source FNA vol. 12, p. 215. FNA vol. 12, p. 219.
Parent taxa Euphorbiaceae > Croton Euphorbiaceae > Croton
Sibling taxa
C. alabamensis, C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
C. alabamensis, C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. glandulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
Subordinate taxa
C. glandulosus var. arenicola, C. glandulosus var. floridanus, C. glandulosus var. lindheimeri, C. glandulosus var. pubentissimus, C. glandulosus var. septentrionalis
Synonyms Decarinium glandulosum, Geiseleria glandulosa, Oxydectes glandulosa C. fergusonii
Name authority Linnaeus: Syst. Nat. ed. 10, 2: 1275. (1759) — (as glandulosum) Jacquin: Enum. Syst. Pl., 32. (1760) — (as lineare)
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