Flora of North America species comparison

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Croton glandulosus

Croton fruticulosus

sand croton, sand or tooth-leaf or Tropic croton, tooth-leaf croton, Tropic croton, vente conmigo

bush croton, encinilla, hierba loca, shrubby croton
Habit Herbs, annual, 1–12 dm, monoecious. Shrubs, 2–10 dm, monoecious.
Stems

much branched distally, usually coarsely stellate-hairy, rarely glabrescent.

much branched distally, stellate-hairy.
Leaves

not clustered;

stipules linear-subulate, glandular or not, to 0.5 mm or absent;

petiole 0.2–1 cm, glands at apex 2, yellow, sessile to shortly stipitate, cuplike;

blade ovate proximally, oblong-lanceolate distally, 0.6–3.5(–7) × 0.3–1.5(–3) cm, base obtuse to truncate, margins coarsely crenate to serrate-dentate, apex obtuse to rounded, both surfaces green, stellate-hairy, glabrate, or rarely glabrous.

not clustered;

stipules rudimentary or absent;

petiole 0.4–1(–1.5) cm, 1/8–1/5 blade length, glands absent at apex;

blade ovate to ovate-lanceolate, 2–8 × 2–4 cm, base truncate to cordate, margins serrulate, often slightly undulate, apex attenuate, acute, abaxial surface pale green, stellate-hairy, adaxial surface darker green, puberulent.
Inflorescences

bisexual, racemes, 1–3 cm, staminate flowers 10–20, pistillate flowers 1–4.

bisexual, racemes, 3–12 cm, staminate flowers 10–20, pistillate flowers 2–5.
Pedicels

staminate 0.8–2 mm, pistillate 0–5 mm.

staminate 2.5–4 mm, pistillate 0–0.5 mm.
Staminate flowers

sepals 5, 0.8–1.2 mm, abaxial surface stellate-hairy;

petals 5, oblanceolate, 1–1.3 mm, abaxial surface glabrous except margins ciliate;

stamens 7–13.

sepals 5, 0.8–1.2 mm, abaxial surface stellate-tomentose;

petals 5, oblanceolate to spatulate, 2 mm, abaxial surface glabrous except margins densely fimbrillate-villous;

stamens 9–16.
Pistillate flowers

sepals 5, subequal, 6–7.5 mm, margins entire, apex straight to slightly incurved, abaxial surface glabrous except stellate-hairy apically;

petals 0 or 5, rudimentary;

ovary 3-locular;

styles 3, 1–2.5 mm, deeply 2-fid, terminal segments 6.

sepals 5, equal, 2.2 mm, margins entire, apex incurved, abaxial surface stellate-hairy;

petals 0;

ovary 3-locular;

styles 3, 3–4.5 mm, 2-fid to base, terminal segments 6.
Capsules

3.5–6 × 4–5 mm, smooth;

columella 3-angled.

5–6 mm diam., smooth;

columella apex with 3 rounded, inflated lobes.
Seeds

3–4 × 2–2.5 mm, shiny.

4–5 × 3.2–3.8 mm, shiny.
2n

= 16.

Croton glandulosus

Croton fruticulosus
Phenology Flowering May–Dec.
Habitat Limestone or basalt hills.
Elevation 100–1700 m. (300–5600 ft.)
Distribution
from FNA
AL; AR; DE; FL; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; NC; NE; NJ; NM; OH; OK; PA; SC; TN; TX; VA; WI; WV; Mexico; Central America; South America; West Indies
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; TX; Mexico (Chihuahua, Coahuila, Nuevo León, San Luis Potosí, Tamaulipas)
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties ca. 20 (5 in the flora).

Croton glandulosus is widespread in the New World, with a complex pattern of variation. The classification here follows B. W. van Ee et al. (2009).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Croton fruticulosus is known in the flora area from southeastern Arizona through southern New Mexico and trans-Pecos Texas to the Edwards Plateau.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Key
1. Leaf blades glabrate or sparsely to moderately stellate-hairy.
→ 2
2. Plants 2–12 dm; leaf blades 2–7 cm.
var. septentrionalis
2. Plants 1–2 dm; leaf blades 0.6–2(–3) cm.
→ 3
3. Leaf blades glabrate, bases markedly 3-veined; petiole apical glands sessile; Florida.
var. floridanus
3. Leaf blades sparsely stellate-hairy, bases obscurely 3-veined; petiole apical glands stipitate; Kansas, New Mexico, Oklahoma, Texas.
var. lindheimeri
1. Leaf blades moderately to densely stellate-hairy.
→ 4
4. Leaf blades firm-thick, length mostly 2 times width or less, marginal teeth rounded; petiole apical glands sessile.
var. arenicola
4. Leaf blades membranous, length mostly more than 2 times width, marginal teeth pointed; petiole apical glands sessile or stipitate.
→ 5
5. Leaf blades 1–2(–3) × 0.3–0.8(–1.3) cm; petiole apical glands stipitate, circular when dry, 0.1–0.4 mm diam.
var. lindheimeri
5. Leaf blades 2–7 × 0.7–3 cm; petiole apical glands sessile, wavy-wrinkled when dry, 0.5–0.8 mm diam.
→ 6
6. Stems densely stellate-hairy, hairs spreading, radii unequal; leaf blades densely stellate-villous.
var. pubentissimus
6. Stems moderately stellate-hairy, hairs appressed, radii equal; leaf blades moderately stellate-hairy.
var. septentrionalis
Source FNA vol. 12, p. 215. FNA vol. 12, p. 214.
Parent taxa Euphorbiaceae > Croton Euphorbiaceae > Croton
Sibling taxa
C. alabamensis, C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
C. alabamensis, C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. glandulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
Subordinate taxa
C. glandulosus var. arenicola, C. glandulosus var. floridanus, C. glandulosus var. lindheimeri, C. glandulosus var. pubentissimus, C. glandulosus var. septentrionalis
Synonyms Decarinium glandulosum, Geiseleria glandulosa, Oxydectes glandulosa
Name authority Linnaeus: Syst. Nat. ed. 10, 2: 1275. (1759) — (as glandulosum) Engelmann ex Torrey: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 194. (1859) — (as fruticulosum)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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