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sand croton, sand or tooth-leaf or Tropic croton, tooth-leaf croton, Tropic croton, vente conmigo

Alabama croton

Habit Herbs, annual, 1–12 dm, monoecious. Shrubs, 5–35 dm, monoecious.
Stems

much branched distally, usually coarsely stellate-hairy, rarely glabrescent.

usually well branched distally, lepidote.

Leaves

not clustered;

stipules linear-subulate, glandular or not, to 0.5 mm or absent;

petiole 0.2–1 cm, glands at apex 2, yellow, sessile to shortly stipitate, cuplike;

blade ovate proximally, oblong-lanceolate distally, 0.6–3.5(–7) × 0.3–1.5(–3) cm, base obtuse to truncate, margins coarsely crenate to serrate-dentate, apex obtuse to rounded, both surfaces green, stellate-hairy, glabrate, or rarely glabrous.

clustered at branch tips;

stipules absent;

petiole 0.6–2 cm, glands absent at apex;

blade elliptic, ovate, or oblong, 3–10 × 1.5–5 cm, base rounded to obtuse, margins entire, sometimes ± undulate, apex acute, rounded, or emarginate, abaxial surface silvery or coppery, densely lepidote, adaxial surface green, sparsely lepidote.

Inflorescences

bisexual, racemes, 1–3 cm, staminate flowers 10–20, pistillate flowers 1–4.

bisexual or unisexual, racemes, 2–4.5 cm, staminate flowers 0–15, pistillate flowers 0–10.

Pedicels

staminate 0.8–2 mm, pistillate 0–5 mm.

staminate 2.2–4 mm, pistillate 2.2–7.5 mm (7–11 mm in fruit).

Staminate flowers

sepals 5, 0.8–1.2 mm, abaxial surface stellate-hairy;

petals 5, oblanceolate, 1–1.3 mm, abaxial surface glabrous except margins ciliate;

stamens 7–13.

sepals 5, 1.1–2.9 mm, abaxial surface lepidote;

petals 5, oblong-ovate, 2–3.1 mm, abaxial surface glabrous except margins stellate-ciliate;

stamens 10–22.

Pistillate flowers

sepals 5, subequal, 6–7.5 mm, margins entire, apex straight to slightly incurved, abaxial surface glabrous except stellate-hairy apically;

petals 0 or 5, rudimentary;

ovary 3-locular;

styles 3, 1–2.5 mm, deeply 2-fid, terminal segments 6.

sepals 5, equal, 2–4.5 mm, margins entire, apex incurved, abaxial surface lepidote;

petals 5, pale green, ovate, 2–3.5 mm;

ovary 3-locular;

styles 3, 2–5 mm, usually unbranched, rarely 2-fid, terminal segments 3 (or 6).

Capsules

3.5–6 × 4–5 mm, smooth;

columella 3-angled.

1.6–2.5 × 2–3 mm, smooth;

columella 3-angled.

Seeds

3–4 × 2–2.5 mm, shiny.

6.7–8 × 5.2–6 mm, shiny.

2n

= 16.

Croton glandulosus

Croton alabamensis

Distribution
from FNA
AL; AR; DE; FL; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; NC; NE; NJ; NM; OH; OK; PA; SC; TN; TX; VA; WI; WV; Mexico; Central America; South America; West Indies
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties ca. 20 (5 in the flora).

Croton glandulosus is widespread in the New World, with a complex pattern of variation. The classification here follows B. W. van Ee et al. (2009).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

B. W. van Ee et al. (2006) examined populations of Croton alabamensis using DNA sequence and AFLP data, and their results supported the recognition of two varieties and an isolated position in the genus. Buds develop in the summer and fall for the following spring's flowers.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades glabrate or sparsely to moderately stellate-hairy.
→ 2
2. Plants 2–12 dm; leaf blades 2–7 cm.
var. septentrionalis
2. Plants 1–2 dm; leaf blades 0.6–2(–3) cm.
→ 3
3. Leaf blades glabrate, bases markedly 3-veined; petiole apical glands sessile; Florida.
var. floridanus
3. Leaf blades sparsely stellate-hairy, bases obscurely 3-veined; petiole apical glands stipitate; Kansas, New Mexico, Oklahoma, Texas.
var. lindheimeri
1. Leaf blades moderately to densely stellate-hairy.
→ 4
4. Leaf blades firm-thick, length mostly 2 times width or less, marginal teeth rounded; petiole apical glands sessile.
var. arenicola
4. Leaf blades membranous, length mostly more than 2 times width, marginal teeth pointed; petiole apical glands sessile or stipitate.
→ 5
5. Leaf blades 1–2(–3) × 0.3–0.8(–1.3) cm; petiole apical glands stipitate, circular when dry, 0.1–0.4 mm diam.
var. lindheimeri
5. Leaf blades 2–7 × 0.7–3 cm; petiole apical glands sessile, wavy-wrinkled when dry, 0.5–0.8 mm diam.
→ 6
6. Stems densely stellate-hairy, hairs spreading, radii unequal; leaf blades densely stellate-villous.
var. pubentissimus
6. Stems moderately stellate-hairy, hairs appressed, radii equal; leaf blades moderately stellate-hairy.
var. septentrionalis
1. Leaf blade abaxial surfaces silvery, scales mostly unpigmented or light amber, some dark blackish brown scales sometimes present; inflorescences 9–18-flowered, producing 0–11 fruits; Alabama.
var. alabamensis
1. Leaf blade abaxial surfaces coppery, some scales unpigmented, others with dark reddish brown center and reddish amber rays; inflorescences 6–14-flowered, producing 0–6 fruits; Texas.
var. texensis
Source FNA vol. 12, p. 215. FNA vol. 12, p. 210.
Parent taxa Euphorbiaceae > Croton Euphorbiaceae > Croton
Sibling taxa
C. alabamensis, C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. glandulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
Subordinate taxa
C. glandulosus var. arenicola, C. glandulosus var. floridanus, C. glandulosus var. lindheimeri, C. glandulosus var. pubentissimus, C. glandulosus var. septentrionalis
C. alabamensis var. alabamensis, C. alabamensis var. texensis
Synonyms Decarinium glandulosum, Geiseleria glandulosa, Oxydectes glandulosa
Name authority Linnaeus: Syst. Nat. ed. 10, 2: 1275. (1759) — (as glandulosum) E. A. Smith ex Chapman: Fl. South. U.S. ed. 2, 648. (1883)
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