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Cortez's croton, palillo

capitate croton, hogwort, woolly croton

Habit Shrubs, 10–30 dm, dioecious. Herbs, annual, 3–8 dm, monoecious.
Stems

di- or trichotomously branched, stellate-hairy, young stems often blackened by fungus.

well branched distally, stellate-hairy, hairs whitish to pale yellow, glabrescent.

Leaves

not clustered;

stipules rudimentary;

petiole 1–1.5 cm, 1/6–1/3 leaf blade length, glands absent at apex;

blade narrowly obovate, oblong, elliptic, or lanceolate, 3–10 × 1–5 cm, mostly less than 3.5 times as long as wide, base subacute to rounded, margins minutely denticulate, apex acute or acuminate, abaxial surface light green, densely ochraceous stellate-hispid when young, pale stellate-hairy when mature, adaxial surface darker green, glabrous.

not clustered;

stipules filiform, 2–5 mm;

petiole 0.5–6 cm, glands absent at apex;

blade ovate to lanceolate-elliptic, 3–8(–15) × 1–4 cm, base rounded to cuneate, margins entire, apex obtuse to acute or rounded, abaxial surface light green, not appearing brown-dotted, no stellate hairs with brown centers, adaxial surface darker green, both appressed stellate-hairy.

Inflorescences

unisexual, racemes;

staminate 7–20 cm, flowers 10–30;

pistillate 1–5 cm, flowers 8–20(–30).

bisexual, racemes or capitate, 1.5–3 cm, staminate flowers 5–15, pistillate flowers 3–6.

Pedicels

staminate 0–1 mm, pistillate 0–0.5 mm.

staminate 0.5–1.5 mm, pistillate 0–1.5 mm.

Staminate flowers

sepals 5, 2.5 mm, abaxial surface stellate-hairy;

petals 5, oblong, 3 mm, abaxial surface glabrous except margins ciliate basally;

stamens 12–16.

sepals 5, 0.8–1 mm, abaxial surface stellate-hairy;

petals 5, linear-oblong-lanceolate, 0.8–1 mm, abaxial surface stellate-hairy;

stamens 7–12.

Pistillate flowers

sepals 5, equal, 1–1.5 mm, margins entire, apex straight to slightly incurved, abaxial surface stellate-hispid;

petals rudimentary;

ovary 3-locular;

styles 3, 4–5 mm, 2-fid, terminal segments 6.

sepals 6–9, equal, 7–10 mm to 15 mm in fruit, opening broadly and persistent after capsule dehiscence, margins entire, apex recurved, abaxial surface stellate-hairy;

petals 0;

ovary 3-locular;

styles 3, 2–3 mm, 2–3 times 2-fid, terminal segments 12–18(–24).

Capsules

5–6 × 4.5–5.5 mm, smooth;

columella apex with 3 rounded, inflated lobes.

7–9 × 5–6 mm, smooth;

columella tipped with 3-pronged grappling hooklike appendage.

Seeds

4–5 × 3–3.5 mm, shiny (with silvery sheen).

5 × 4.5–5 mm, shiny.

2n

= 20.

= 20.

Croton cortesianus

Croton capitatus

Phenology Flowering Feb–Oct, possibly year-round. Flowering Jun–Oct.
Habitat Scrub forests. Pastures, old fields, cultivated land, prairies, floodplains, longleaf pinelands, sandy to loamy soils.
Elevation 0–50 m. (0–200 ft.) 0–300 m. (0–1000 ft.)
Distribution
from FNA
TX; e Mexico; se Mexico; Central America
[BONAP county map]
from FNA
AL; AR; DC; DE; IA; IL; IN; KS; KY; MA; MD; MO; MS; NC; NE; NJ; OH; OK; PA; SC; TN; TX; VA; WV
[WildflowerSearch map]
[BONAP county map]
Discussion

Croton cortesianus is known in the flora area only from Cameron, Hidalgo, and Starr counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Croton capitatus is frequently confused with C. heptalon and C. lindheimeri. Croton capitatus usually has some leaves that are blunt-tipped and mucronate (versus all or most acute in C. lindheimeri), and the pistillate sepals are longer than the ovary, with tips that flare outward at anthesis. Also, C. capitatus has petioles roughly equal in length from the middle to the tip of the stem, whereas C. lindheimeri has petioles that decrease more markedly in length from the middle to the tip of the stem. Occurrences of C. capitatus east of the Appalachian Mountains are likely adventive.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 213. FNA vol. 12, p. 212.
Parent taxa Euphorbiaceae > Croton Euphorbiaceae > Croton
Sibling taxa
C. alabamensis, C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. glandulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
C. alabamensis, C. argenteus, C. argyranthemus, C. californicus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. glandulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
Synonyms C. chichenensis, C. trichocarpus Heptallon graveolens, Oxydectes capitata, Pilinophytum capitatum
Name authority Kunth: in A. von Humboldt et al., Nov. Gen. Sp. 2(fol.): 66; 2(qto.): 83. (1817) Michaux: Fl. Bor.-Amer. 2: 214. (1803) — (as capitatum)
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