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Alabama croton

grassland croton, hierba del gato, rosval, rubaldo

Habit Shrubs, 5–35 dm, monoecious. Subshrubs, 2–5(–9) dm, dioecious.
Stems

usually well branched distally, lepidote.

well-branched from base, stellate-lepidote.

Leaves

clustered at branch tips;

stipules absent;

petiole 0.6–2 cm, glands absent at apex;

blade elliptic, ovate, or oblong, 3–10 × 1.5–5 cm, base rounded to obtuse, margins entire, sometimes ± undulate, apex acute, rounded, or emarginate, abaxial surface silvery or coppery, densely lepidote, adaxial surface green, sparsely lepidote.

not clustered;

stipules rudimentary or absent;

petiole 0.2–0.8(–2) cm, usually less than 1/2 blade length, glands absent at apex;

blade narrowly elliptic-ovate to lanceolate, 1–6.5 × 0.6–2.2 cm, usually less than 1/2 as wide as long, base rounded, margins entire, apex acute to rounded, abaxial surface pale green, densely silvery lepidote or stellate-lepidote, adaxial surface darker green, less densely lepidote.

Inflorescences

bisexual or unisexual, racemes, 2–4.5 cm, staminate flowers 0–15, pistillate flowers 0–10.

unisexual, racemes;

staminate 2–8 cm, flowers 4–16;

pistillate 0.5–1 cm, flowers 2–5.

Pedicels

staminate 2.2–4 mm, pistillate 2.2–7.5 mm (7–11 mm in fruit).

staminate 1–4 mm, pistillate 2–5 mm.

Staminate flowers

sepals 5, 1.1–2.9 mm, abaxial surface lepidote;

petals 5, oblong-ovate, 2–3.1 mm, abaxial surface glabrous except margins stellate-ciliate;

stamens 10–22.

sepals 5, 1 mm, abaxial surface lepidote;

petals 0;

stamens 10–12.

Pistillate flowers

sepals 5, equal, 2–4.5 mm, margins entire, apex incurved, abaxial surface lepidote;

petals 5, pale green, ovate, 2–3.5 mm;

ovary 3-locular;

styles 3, 2–5 mm, usually unbranched, rarely 2-fid, terminal segments 3 (or 6).

sepals 5, equal, 1.5–2 mm, margins entire, apex straight to slightly incurved, abaxial surface stellate-lepidote;

petals 0;

ovary 3-locular;

styles 3, 0.5–1.5 mm, 2–3 times 2-fid, terminal segments 12–24.

Capsules

1.6–2.5 × 2–3 mm, smooth;

columella 3-angled.

5–6 mm diam., smooth;

columella 3-winged.

Seeds

6.7–8 × 5.2–6 mm, shiny.

3.5–5 × 3–4 mm, shiny or dull.

2n

= 28, 56.

Croton alabamensis

Croton dioicus

Phenology Flowering Mar–Nov.
Habitat Limestone and igneous mountains, canyons, mesas, flats, disturbed areas.
Elevation 30–2000 m. (100–6600 ft.)
Distribution
from FNA
AL; TX
[WildflowerSearch map]
[BONAP county map]
from FNA
NM; TX; Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

B. W. van Ee et al. (2006) examined populations of Croton alabamensis using DNA sequence and AFLP data, and their results supported the recognition of two varieties and an isolated position in the genus. Buds develop in the summer and fall for the following spring's flowers.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blade abaxial surfaces silvery, scales mostly unpigmented or light amber, some dark blackish brown scales sometimes present; inflorescences 9–18-flowered, producing 0–11 fruits; Alabama.
var. alabamensis
1. Leaf blade abaxial surfaces coppery, some scales unpigmented, others with dark reddish brown center and reddish amber rays; inflorescences 6–14-flowered, producing 0–6 fruits; Texas.
var. texensis
Source FNA vol. 12, p. 210. FNA vol. 12, p. 214.
Parent taxa Euphorbiaceae > Croton Euphorbiaceae > Croton
Sibling taxa
C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. glandulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
C. alabamensis, C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. elliottii, C. fruticulosus, C. glandulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
Subordinate taxa
C. alabamensis var. alabamensis, C. alabamensis var. texensis
Synonyms C. elaeagnifolius, C. gracilis, C. neomexicanus, C. vulpinus
Name authority E. A. Smith ex Chapman: Fl. South. U.S. ed. 2, 648. (1883) Cavanilles: Icon. 1: 4, plate 6. (1791) — (as dioicum)
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