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Alabama croton

California croton, desert croton

Habit Shrubs, 5–35 dm, monoecious. Subshrubs or shrubs, 4–11 dm, dioecious.
Stems

usually well branched distally, lepidote.

loosely branched, stellate-lepidote.

Leaves

clustered at branch tips;

stipules absent;

petiole 0.6–2 cm, glands absent at apex;

blade elliptic, ovate, or oblong, 3–10 × 1.5–5 cm, base rounded to obtuse, margins entire, sometimes ± undulate, apex acute, rounded, or emarginate, abaxial surface silvery or coppery, densely lepidote, adaxial surface green, sparsely lepidote.

not clustered;

stipules absent;

petiole 1–3.5(–4.5) cm, usually less than 1/2 blade length, glands absent at apex;

blade elliptic to narrowly oblong, 2–5.5(–7) × 0.8–2(–2.5) cm, usually more than 2 times as long as wide, base obtuse, margins entire, apex obtuse to rounded, abaxial surface light green, adaxial surface darker green, both stellate-lepidote.

Inflorescences

bisexual or unisexual, racemes, 2–4.5 cm, staminate flowers 0–15, pistillate flowers 0–10.

unisexual, racemes or thyrses;

staminate 0.5–3.5(–10) cm, flowers 3–8(–20);

pistillate 0.5–1 cm, flowers 1–6.

Pedicels

staminate 2.2–4 mm, pistillate 2.2–7.5 mm (7–11 mm in fruit).

staminate 1–5.5(–7) mm, pistillate to 1 mm (1–3 mm in fruit).

Staminate flowers

sepals 5, 1.1–2.9 mm, abaxial surface lepidote;

petals 5, oblong-ovate, 2–3.1 mm, abaxial surface glabrous except margins stellate-ciliate;

stamens 10–22.

sepals 5, 1 mm, abaxial surface stellate-lepidote;

petals 0;

stamens 10–15.

Pistillate flowers

sepals 5, equal, 2–4.5 mm, margins entire, apex incurved, abaxial surface lepidote;

petals 5, pale green, ovate, 2–3.5 mm;

ovary 3-locular;

styles 3, 2–5 mm, usually unbranched, rarely 2-fid, terminal segments 3 (or 6).

sepals 5, equal, 2 mm, margins entire, apex incurved, abaxial surface stellate-lepidote;

petals 0;

ovary 3-locular;

styles 3, 2–2.5 mm, 4-fid, terminal segments usually 12.

Capsules

1.6–2.5 × 2–3 mm, smooth;

columella 3-angled.

6–8 × 5–7 mm, smooth;

columella 3-winged.

Seeds

6.7–8 × 5.2–6 mm, shiny.

4–5.5 × 3.5–5.5 mm, dull.

2n

= 28.

Croton alabamensis

Croton californicus

Phenology Flowering Feb–Oct.
Habitat Sandy soils, sage scrub, dunes, washes.
Elevation 0–900 m. (0–3000 ft.)
Distribution
from FNA
AL; TX
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NV; UT; Mexico (Baja California, Sinaloa)
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

B. W. van Ee et al. (2006) examined populations of Croton alabamensis using DNA sequence and AFLP data, and their results supported the recognition of two varieties and an isolated position in the genus. Buds develop in the summer and fall for the following spring's flowers.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blade abaxial surfaces silvery, scales mostly unpigmented or light amber, some dark blackish brown scales sometimes present; inflorescences 9–18-flowered, producing 0–11 fruits; Alabama.
var. alabamensis
1. Leaf blade abaxial surfaces coppery, some scales unpigmented, others with dark reddish brown center and reddish amber rays; inflorescences 6–14-flowered, producing 0–6 fruits; Texas.
var. texensis
Source FNA vol. 12, p. 210. FNA vol. 12, p. 211.
Parent taxa Euphorbiaceae > Croton Euphorbiaceae > Croton
Sibling taxa
C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. glandulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
C. alabamensis, C. argenteus, C. argyranthemus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. glandulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
Subordinate taxa
C. alabamensis var. alabamensis, C. alabamensis var. texensis
Synonyms Hendecandra procumbens, C. californicus var. longipes, C. californicus var. mohavensis, C. californicus var. tenuis, C. longipes, C. mohavensis, C. tenuis
Name authority E. A. Smith ex Chapman: Fl. South. U.S. ed. 2, 648. (1883) Müller Arg.: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 15(2): 691. (1866)
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