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Alabama croton

healing croton, silver croton, silverleaf croton

Habit Shrubs, 5–35 dm, monoecious. Herbs or subshrubs, annual or short-lived perennial, 1–6 dm, monoecious.
Stems

usually well branched distally, lepidote.

several from base, branching distally into 2–4 branches, lepidote.

Leaves

clustered at branch tips;

stipules absent;

petiole 0.6–2 cm, glands absent at apex;

blade elliptic, ovate, or oblong, 3–10 × 1.5–5 cm, base rounded to obtuse, margins entire, sometimes ± undulate, apex acute, rounded, or emarginate, abaxial surface silvery or coppery, densely lepidote, adaxial surface green, sparsely lepidote.

not clustered;

stipules subulate, 0.2 mm;

petiole 0.2–1(–2) cm, glands absent at apex;

blade on proximal parts of stems oval to narrowly obovate, on distal parts oblong to lanceolate-oblong or broadly elliptic, 1–5 × 0.5–2(–3) cm, base rounded to narrowed, margins entire, apex obtuse to rounded, abaxial surface silvery, densely lepidote, adaxial surface darker green, sparsely lepidote or glabrate.

Inflorescences

bisexual or unisexual, racemes, 2–4.5 cm, staminate flowers 0–15, pistillate flowers 0–10.

bisexual, racemes, 2–5 cm, staminate flowers 15–35, pistillate flowers 2–8.

Pedicels

staminate 2.2–4 mm, pistillate 2.2–7.5 mm (7–11 mm in fruit).

staminate 1–5 mm, pistillate 0–2 mm.

Staminate flowers

sepals 5, 1.1–2.9 mm, abaxial surface lepidote;

petals 5, oblong-ovate, 2–3.1 mm, abaxial surface glabrous except margins stellate-ciliate;

stamens 10–22.

sepals 5, 5 mm, abaxial surface lepidote;

petals 5, oblong-spatulate, 5 mm, abaxial surface densely lepidote, scales translucent, petals appearing hyaline;

stamens 10–15.

Pistillate flowers

sepals 5, equal, 2–4.5 mm, margins entire, apex incurved, abaxial surface lepidote;

petals 5, pale green, ovate, 2–3.5 mm;

ovary 3-locular;

styles 3, 2–5 mm, usually unbranched, rarely 2-fid, terminal segments 3 (or 6).

sepals 5–7, connate for 1/2+ length, equal, 3–4 mm, margins entire, apex incurved, abaxial surface lepidote;

petals 0;

ovary 3-locular;

styles 3, 2–4 mm, 1–3 times 2-fid at apex, terminal segments 6–24.

Capsules

1.6–2.5 × 2–3 mm, smooth;

columella 3-angled.

5–6 × 3–4 mm, smooth;

columella 3-angled.

Seeds

6.7–8 × 5.2–6 mm, shiny.

4–5 × 2.5–3 mm, dull.

Croton alabamensis

Croton argyranthemus

Phenology Flowering Mar–Sep.
Habitat Deep sandy soils in pinelands, pine-oak scrub, sandhills.
Elevation 0–200 m. (0–700 ft.)
Distribution
from FNA
AL; TX
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; FL; GA; LA; MS; TX; Mexico (Nuevo León, Tamaulipas)
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

B. W. van Ee et al. (2006) examined populations of Croton alabamensis using DNA sequence and AFLP data, and their results supported the recognition of two varieties and an isolated position in the genus. Buds develop in the summer and fall for the following spring's flowers.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Croton argyranthemus is closely related to C. coryi. Although they have very different kinds of vegetative indumentum, the staminate petals are characteristically silvery-lepidote in both species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blade abaxial surfaces silvery, scales mostly unpigmented or light amber, some dark blackish brown scales sometimes present; inflorescences 9–18-flowered, producing 0–11 fruits; Alabama.
var. alabamensis
1. Leaf blade abaxial surfaces coppery, some scales unpigmented, others with dark reddish brown center and reddish amber rays; inflorescences 6–14-flowered, producing 0–6 fruits; Texas.
var. texensis
Source FNA vol. 12, p. 210. FNA vol. 12, p. 211.
Parent taxa Euphorbiaceae > Croton Euphorbiaceae > Croton
Sibling taxa
C. argenteus, C. argyranthemus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. glandulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
C. alabamensis, C. argenteus, C. californicus, C. capitatus, C. ciliatoglandulifer, C. cortesianus, C. coryi, C. dioicus, C. elliottii, C. fruticulosus, C. glandulosus, C. heptalon, C. humilis, C. incanus, C. leucophyllus, C. lindheimeri, C. lindheimerianus, C. linearis, C. michauxii, C. monanthogynus, C. parksii, C. pottsii, C. punctatus, C. sancti-lazari, C. setigerus, C. soliman, C. sonorae, C. suaveolens, C. texensis, C. wigginsii
Subordinate taxa
C. alabamensis var. alabamensis, C. alabamensis var. texensis
Name authority E. A. Smith ex Chapman: Fl. South. U.S. ed. 2, 648. (1883) Michaux: Fl. Bor.-Amer. 2: 215. (1803) — (as argyranthemum)
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