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beak hawk's-beard, weedy hawksbeard

gray hawksbeard, largeflower hawksbeard, western hawk's beard

Habit Annuals, biennials, or perennials, 3–120 cm (taproots slender to thick, caudices swollen). Perennials, 8–40 cm; taproots deep, caudices swollen, (often covered with old leaf bases).
Stems

1, erect to arcuate or decumbent (green or purple proximally), usually much branched, glabrate to hispid and/or tomentose, sometimes sparsely setose (setae black).

1–3, erect, stout, branched from bases or beyond, hispid, tomentose, or tomentulose, sometimes stipitate-glandular distally.

Leaves

basal and cauline;

petiolate (bases clasping);

blades oblanceolate to ovate, often runcinate, 10–35 × 2–8 cm, margins pinnately lobed to toothed (terminal lobes relatively large), apices obtuse or acute, faces usually hirsute (hairs sometimes only on veins) or glabrous (cauline sessile, bases auriculate, clasping, margins ± toothed).

basal and cauline;

petiolate;

blades elliptic, runcinate, (5–)8–20 × 2–5 cm, margins pinnately-lobed to sinuously dentate (lobes broadly lanceolate, often dentate), apices acute or acuminate, faces gray-tomentose, sometimes stipitate-glandular.

Involucres

cylindro-campanulate (becoming turbinate or urceolate in fruit), 5–14 × 5–6 mm.

cylindric, 11–19 × 5–10 mm.

Florets

50–70;

corollas yellow (reddish abaxially), 6–15 mm.

10–40;

corollas yellow, 18–22 mm.

Phyllaries

7–16, (reflexed at maturity) lanceolate, 10–12 mm, (margins green to yellowish), apices obtuse or acute (ciliate), abaxial faces tomentose and often stipitate-glandular, adaxial with fine, appressed hairs.

7–13, lanceolate, 12–15 mm, (bases thickened, keeled, margins green, often scarious) apices acute or acuminate, abaxial faces gray-tomentose, sometimes setose (setae black or greenish) or stipitate-glandular, adaxial glabrous or with fine hairs.

Calyculi

of 5–12, ovate to linear-lanceolate, glabrous bractlets 3–4 mm (reflexed in fruit, scarious).

of 6–8, lanceolate or linear, glabrate to tomentose bractlets 2–6 mm.

Heads

10–20, in lax, corymbiform arrays.

2–30, in loose corymbiform arrays.

Cypselae

(monomorphic or dimorphic) pale brown or yellowish, fusiform, 4–9 mm, outer wider with apices attenuate (not beaked), inner gradually tapered, beaked (beaks 2–5 mm, ± equal to bodies), ribs 10 (narrow);

pappi white (fine, soft), 3–6 mm.

golden or dark brown, subcylindric, 6–10 mm, apices tapered (not beaked), ribs 10–18, strong and rounded;

pappi yellowish white, 10–12 mm (bristles unequal).

2n

= 8, 16.

= 22, 33, 44, 55, 66, 77, 88.

Crepis vesicaria

Crepis occidentalis

Phenology Flowering Feb–Oct.
Habitat Sandy clearings, hillsides
Elevation 0–300 m (0–1000 ft)
Distribution
from FNA
CA; CT; NC; NY; OR; PA; BC; Europe [Introduced in North America; introduced, South America]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; NM; NV; OR; SD; UT; WA; WY; AB; BC; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Native to the Mediterranean region of western Europe, Crepis vesicaria is recognized by its annual or biennial habit, pinnately lobed leaves, reflexed calyculi, tomentose and glandular phyllaries, and slender, long-beaked inner cypselae. It is polymorphic; subspecies are recognized in Europe. E. B. Babcock (1947) identified the North American plants as subsp. taraxaciflora (Thuiller) Thellung, which some Europeans (T. G. Tutin et al. 1964–1980, vol. 4) have listed as a synonym of subsp. haenseleri (Boissier ex de Candolle) P. D. Sell.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 4 (4 in the flora).

Crepis occidentalis is recognized by the old, brown leaf bases persisting on caudices, by stems, leaves, and phyllaries gray-tomentose, and by loose, corymbiform arrays with relatively few, relatively large heads. It is widespread and polymorphic. Some specimens have coarse setae or black, stipitate glands on the phyllaries in addition to the tomentose indument, the stipitate glands sometimes extending proximally on stems. Four intergrading subspecies were recognized by E. B. Babcock (1947). The sexual diploid forms are found in subsp. occidentalis and occur in northern California and adjacent Nevada. The other subspecies are polyploid and apomictic (Babcock).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Phyllaries sparsely to densely stipitate-glandular
→ 2
1. Phyllaries 8–12, usually eglandular, if glandular, phyllaries 8
→ 3
2. Phyllaries (peduncles and distal cauline leaves) stipitate-glandular (lacking large dark or black glandular setae); phyllaries 7–8 or 10–13; florets 18–30
subsp. occidentalis
2. Phyllaries (peduncles and distal cauline leaves) stipitate-glandular (and with dark or black, glandular setae); phyllaries 8, florets 10–14
subsp. costata
3. Plants 10–40 cm (stems with definite primary axes, branched distally; phyllaries mostly 8; leaves coarsely dentate or pinnately lobed (lobes closely spaced)
subsp. pumila
3. Plants 5–20 cm (stems branched proximally; phyllaries 8–12; leaves deeply pinnately lobed (lobes remotely spaced, lanceolate, or linear, entire or dentate)
subsp. conjuncta
Source FNA vol. 19, p. 238. FNA vol. 19, p. 232.
Parent taxa Asteraceae > tribe Cichorieae > Crepis Asteraceae > tribe Cichorieae > Crepis
Sibling taxa
C. acuminata, C. atribarba, C. bakeri, C. barbigera, C. biennis, C. bursifolia, C. capillaris, C. elegans, C. foetida, C. intermedia, C. modocensis, C. monticola, C. nana, C. nicaeënsis, C. occidentalis, C. pannonica, C. pleurocarpa, C. pulchra, C. rubra, C. runcinata, C. setosa, C. tectorum, C. zacintha
C. acuminata, C. atribarba, C. bakeri, C. barbigera, C. biennis, C. bursifolia, C. capillaris, C. elegans, C. foetida, C. intermedia, C. modocensis, C. monticola, C. nana, C. nicaeënsis, C. pannonica, C. pleurocarpa, C. pulchra, C. rubra, C. runcinata, C. setosa, C. tectorum, C. vesicaria, C. zacintha
Subordinate taxa
C. occidentalis subsp. conjuncta, C. occidentalis subsp. costata, C. occidentalis subsp. occidentalis, C. occidentalis subsp. pumila
Synonyms Psilochenia occidentalis
Name authority Linnaeus: Sp. Pl. 2: 805. (1753) Nuttall: J. Acad. Nat. Sci. Philadelphia 7: 29. (1834)
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