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mountain river hawthorn, river hawthorn
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| Shrubs or trees, 30–50 dm. |
Shrubs or trees, 60 dm, branches ± weeping. |
twigs: new growth greenish, sparsely pubescent, early glabrescent, 1–2-years old often red-purple; bark on younger 2–5 cm thick branches dark gray-brown, sometimes copper-colored; thorns on twigs straight or ± recurved, 2-years old black or purple-black, glossy, fine, 1.5–4 cm. |
twigs: new growth green, pubescent, 1-year old purplish brown under exfoliating wax, older deep dull gray; thorns on twigs absent or numerous, straight to slightly recurved, 1-year old often copper to deep red, 1.5–5 cm. |
petiole 1–2.5 cm, glabrous; blade elliptic to narrowly elliptic, 3–8 cm, length 2+ times width, thin, base cuneate, lobes 0 or small apiculi at ends of some vein tips, margins serrate, teeth numerous, usually acute, fine to coarse, venation semicamptodromous, veins 4 or 5 per side, larger often branched proximal to ends, apex acute, abaxial surface glabrous, adaxial appressed-strigose young. |
petiole length 30–40% blade, roughly pubescent, black-glandular; blade dull green, abaxially paler, oblong-spatulate to cuneate or narrowly obdeltate, 3 cm, thin, ± floppy, base cuneate, lobes 1 or 2 per side distally, well defined, lobe apex acute, margins strongly crenate-serrate almost to bases, densely glandular, glands black, veins (2 or)3 or 4(or 5) per side, apex usually cuspidate, sometimes acute, adaxial surface sparsely pubescent young, main veins and axils hairy; on extension shoots larger, relatively wider, more deeply incised. |
6–12-flowered; branches glandular-punctate; bracteoles early caducous, usually present, margins sessile-glandular. |
3–7-flowered; branches densely short-canescent; bracteoles deciduous, linear, margins short-stipitate-glandular, adaxially short-pubescent. |
14–17 mm diam.; sepals broadly deltate, 6–8 mm, margins entire, apex narrow, glabrous; stamens 10, anthers pink, sometimes ivory; styles 3–5. |
15–20 mm diam.; hypanthium tomentose; sepals narrowly triangular, margins glandular-serrate, abaxially pubescent; anther color not recorded; styles 3–5. |
deep red (mid Aug), black or blackish purple mature, suborbicular, 10 mm diam.; sepals ± erect to strongly outcurved or ± patent, 2–3 mm; pyrenes 3 or 4, sides eroded. |
1–3 per infructescence, deep red, suborbicular, 10 mm diam., punctate, glabrous; sepals patent to reflexed; pyrenes 3–5. |
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| Flowering mid May–early Jun; fruiting Aug–Sep. |
Flowering Apr–early May; fruiting Aug–Sep. |
| Intermontane streamsides, ditches, flood plains |
Dry pine woods, open scrub, sand plains |
| 1300–2300 m (4300–7500 ft) |
100–800 m (300–2600 ft) |
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AZ; CO; ID; NM; NV; UT; WY
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AL; FL; GA; LA; NC; SC |
Crataegus rivularis is most abundant in intermontane Colorado and Utah and is one of two hawthorns found in Arizona; it is rare in Nevada. There are reports from northwestern Texas (for example, D. S. Correll and M. C. Johnston 1970); they have not been verified. Crataegus rivularis is one of the more common woody species of mesic, valley bottom intermontane habitats. The species flowers early. Crataegus rivularis is superficially similar to the more or less sympatric, less common, C. saligna; it has larger leaves with fewer veins, larger flowers with ten stamens, and larger pomes. It differs from C. erythropoda little other than in leaf shape and ripe fruit color. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Crataegus senta is widespread from western North Carolina and South Carolina (where it is most common) to northern Florida and Georgia, with a disjunct record from West Feliciana Parish, Louisiana. Habitats are typical for the series. Crataegus senta most resembles C. dispar but is more often confused with C. lancei. Short-shoot leaves of C. senta are narrower and are not as lanate as in C. dispar. Smaller leaves of C. senta have fewer veins (1 or 2 per side); their shape is more like that of C. lancei, but the differences of lobes and leaf teeth are striking (see key and C. lancei discussion). In C. senta, extension-shoot leaves differ little from those of the short shoots, having multiple, relatively short, acute lobes across the subterminal part. By contrast, the extension-shoot leaves of C. lancei are of a long-petiolate, narrowly rhombic form with 1–3 acute to subacute lobes per side. The large, red, cherrylike fruit, similar to that of C. lancei, is very striking. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 9, p. 510. |
FNA vol. 9, p. 631. |
| Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Douglasia > ser. Cerrones |
Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Lacrimatae |
C. aemula, C. aestivalis, C. alabamensis, C. alleghaniensis, C. annosa, C. aprica, C. aquacervensis, C. ashei, C. atrovirens, C. attrita, C. austromontana, C. beata, C. berberifolia, C. biltmoreana, C. brachyacantha, C. brainerdii, C. brazoria, C. brittonii, C. buckleyi, C. calpodendron, C. castlegarensis, C. chrysocarpa, C. coccinea, C. coccinioides, C. cognata, C. collina, C. colonica, C. communis, C. compacta, C. condigna, C. craytonii, C. crocea, C. crus-galli, C. cupressocollina, C. delawarensis, C. dispar, C. dodgei, C. douglasii, C. egens, C. egregia, C. enderbyensis, C. erythropoda, C. exilis, C. eximia, C. extraria, C. fecunda, C. flabellata, C. flava, C. florens, C. floridana, C. florifera, C. fluviatilis, C. formosa, C. frugiferens, C. furtiva, C. gattingeri, C. gaylussacia, C. gilva, C. greggiana, C. harbisonii, C. holmesiana, C. ignava, C. incilis, C. integra, C. intricata, C. invicta, C. iracunda, C. irrasa, C. jesupii, C. jonesiae, C. lacrimata, C. laevigata, C. lanata, C. lancei, C. lanuginosa, C. lassa, C. laurentiana, C. leonensis, C. lepida, C. levis, C. lumaria, C. macracantha, C. macrosperma, C. magniflora, C. margarettae, C. marshallii, C. mendosa, C. meridiana, C. mira, C. mollis, C. monogyna, C. munda, C. nananixonii, C. neobushii, C. nitida, C. oakesiana, C. okanaganensis, C. okennonii, C. opaca, C. opima, C. orbicularis, C. ouachitensis, C. padifolia, C. pennsylvanica, C. persimilis, C. pexa, C. phaenopyrum, C. phippsii, C. pinetorum, C. populnea, C. prona, C. pruinosa, C. pulcherrima, C. punctata, C. purpurella, C. quaesita, C. reverchonii, C. rivuloadamensis, C. rivulopugnensis, C. roribacca, C. rubella, C. rubribracteolata, C. saligna, C. sargentii, C. scabrida, C. schizophylla, C. schuettei, C. segnis, C. senta, C. sheila-phippsiae, C. sheridana, C. shuswapensis, C. sororia, C. spathulata, C. spes-aestatum, C. stolonifera, C. stonei, C. submollis, C. suborbiculata, C. succulenta, C. tecta, C. teres, C. texana, C. tracyi, C. triflora, C. turnerorum, C. uniflora, C. ursopedensis, C. venusta, C. viridis, C. visenda, C. wattiana, C. williamsii, C. wootoniana, C. ×atrorubens, C. ×bicknellii, C. ×coleae, C. ×collicola, C. ×disperma, C. ×dispessa, C. ×fretalis, C. ×incaedua, C. ×kelloggii, C. ×latebrosa, C. ×lucorum, C. ×rufula, C. ×sicca, C. ×vailiae |
C. aemula, C. aestivalis, C. alabamensis, C. alleghaniensis, C. annosa, C. aprica, C. aquacervensis, C. ashei, C. atrovirens, C. attrita, C. austromontana, C. beata, C. berberifolia, C. biltmoreana, C. brachyacantha, C. brainerdii, C. brazoria, C. brittonii, C. buckleyi, C. calpodendron, C. castlegarensis, C. chrysocarpa, C. coccinea, C. coccinioides, C. cognata, C. collina, C. colonica, C. communis, C. compacta, C. condigna, C. craytonii, C. crocea, C. crus-galli, C. cupressocollina, C. delawarensis, C. dispar, C. dodgei, C. douglasii, C. egens, C. egregia, C. enderbyensis, C. erythropoda, C. exilis, C. eximia, C. extraria, C. fecunda, C. flabellata, C. flava, C. florens, C. floridana, C. florifera, C. fluviatilis, C. formosa, C. frugiferens, C. furtiva, C. gattingeri, C. gaylussacia, C. gilva, C. greggiana, C. harbisonii, C. holmesiana, C. ignava, C. incilis, C. integra, C. intricata, C. invicta, C. iracunda, C. irrasa, C. jesupii, C. jonesiae, C. lacrimata, C. laevigata, C. lanata, C. lancei, C. lanuginosa, C. lassa, C. laurentiana, C. leonensis, C. lepida, C. levis, C. lumaria, C. macracantha, C. macrosperma, C. magniflora, C. margarettae, C. marshallii, C. mendosa, C. meridiana, C. mira, C. mollis, C. monogyna, C. munda, C. nananixonii, C. neobushii, C. nitida, C. oakesiana, C. okanaganensis, C. okennonii, C. opaca, C. opima, C. orbicularis, C. ouachitensis, C. padifolia, C. pennsylvanica, C. persimilis, C. pexa, C. phaenopyrum, C. phippsii, C. pinetorum, C. populnea, C. prona, C. pruinosa, C. pulcherrima, C. punctata, C. purpurella, C. quaesita, C. reverchonii, C. rivularis, C. rivuloadamensis, C. rivulopugnensis, C. roribacca, C. rubella, C. rubribracteolata, C. saligna, C. sargentii, C. scabrida, C. schizophylla, C. schuettei, C. segnis, C. sheila-phippsiae, C. sheridana, C. shuswapensis, C. sororia, C. spathulata, C. spes-aestatum, C. stolonifera, C. stonei, C. submollis, C. suborbiculata, C. succulenta, C. tecta, C. teres, C. texana, C. tracyi, C. triflora, C. turnerorum, C. uniflora, C. ursopedensis, C. venusta, C. viridis, C. visenda, C. wattiana, C. williamsii, C. wootoniana, C. ×atrorubens, C. ×bicknellii, C. ×coleae, C. ×collicola, C. ×disperma, C. ×dispessa, C. ×fretalis, C. ×incaedua, C. ×kelloggii, C. ×latebrosa, C. ×lucorum, C. ×rufula, C. ×sicca, C. ×vailiae |
| Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 464. (1840) |
Beadle: Bot. Gaz. 30: 341. (1900) |
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