Crataegus rivularis |
Crataegus pruinosa |
|||||||||||||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
mountain river hawthorn, river hawthorn |
frosted hawthorn, waxyfruit hawthorn |
|||||||||||||||||||||
Habit | Shrubs or trees, 30–50 dm. | Shrubs or trees, dense, often suberect, 20–70 dm. | ||||||||||||||||||||
Stems | twigs: new growth greenish, sparsely pubescent, early glabrescent, 1–2-years old often red-purple; bark on younger 2–5 cm thick branches dark gray-brown, sometimes copper-colored; thorns on twigs straight or ± recurved, 2-years old black or purple-black, glossy, fine, 1.5–4 cm. |
compound thorns on trunks present; twigs: new growth reddish brown, 1-year old dull purple-brown, 2-years old dull gray, older paler; thorns on twigs straight to slightly recurved, 2-years old deep purple to shiny black, usually ± fine, 3–5 cm. |
||||||||||||||||||||
Leaves | petiole 1–2.5 cm, glabrous; blade elliptic to narrowly elliptic, 3–8 cm, length 2+ times width, thin, base cuneate, lobes 0 or small apiculi at ends of some vein tips, margins serrate, teeth numerous, usually acute, fine to coarse, venation semicamptodromous, veins 4 or 5 per side, larger often branched proximal to ends, apex acute, abaxial surface glabrous, adaxial appressed-strigose young. |
petiole length 50–66% blade, sparsely glandular; blade trullate to ovate, ovate-oblong, or broadly rhombic, rarely ± deltate, 2–6(–7) cm, subcoriaceous, base broadly cuneate to subtruncate to weakly subcordate, lobes (1–)3 or 4 per side, sinuses shallow to deeper, lobe apex acute, sometimes ± obtuse, margins serrate, veins 5 or 6 per side (except smaller leaves), apex acute, surfaces glabrous (except var. virella). |
||||||||||||||||||||
Inflorescences | 6–12-flowered; branches glandular-punctate; bracteoles early caducous, usually present, margins sessile-glandular. |
5–10-flowered; branches glabrous, sometimes sparsely villous; bracteoles caducous, usually few, margins short-stipitate-glandular. |
||||||||||||||||||||
Flowers | 14–17 mm diam.; sepals broadly deltate, 6–8 mm, margins entire, apex narrow, glabrous; stamens 10, anthers pink, sometimes ivory; styles 3–5. |
15–25 mm diam.; sepals narrowly triangular, 5–6 mm, margins usually entire or subentire, rarely glandular-serrate, abaxially glabrous; stamens (10 or)20, anthers pale pink to bright rose or dull purple, sometimes cream, 0.6–0.8 mm; styles 3–5. |
||||||||||||||||||||
Pomes | deep red (mid Aug), black or blackish purple mature, suborbicular, 10 mm diam.; sepals ± erect to strongly outcurved or ± patent, 2–3 mm; pyrenes 3 or 4, sides eroded. |
greenish with pink or mauve areas, sometimes bright crimson or scarlet, often rather angular, 10–20 mm diam., highly pruinose, not punctate; flesh hard; sepals on collar, spreading; pyrenes 3–5. |
||||||||||||||||||||
Crataegus rivularis |
Crataegus pruinosa |
|||||||||||||||||||||
Phenology | Flowering mid May–early Jun; fruiting Aug–Sep. | |||||||||||||||||||||
Habitat | Intermontane streamsides, ditches, flood plains | |||||||||||||||||||||
Elevation | 1300–2300 m (4300–7500 ft) | |||||||||||||||||||||
Distribution |
AZ; CO; ID; NM; NV; UT; WY
|
AR; CT; GA; IA; IL; IN; KS; KY; MA; ME; MI; MO; NC; NH; NJ; NY; OH; PA; TN; VA; VT; WI; WV; ON; QC
|
||||||||||||||||||||
Discussion | Crataegus rivularis is most abundant in intermontane Colorado and Utah and is one of two hawthorns found in Arizona; it is rare in Nevada. There are reports from northwestern Texas (for example, D. S. Correll and M. C. Johnston 1970); they have not been verified. Crataegus rivularis is one of the more common woody species of mesic, valley bottom intermontane habitats. The species flowers early. Crataegus rivularis is superficially similar to the more or less sympatric, less common, C. saligna; it has larger leaves with fewer veins, larger flowers with ten stamens, and larger pomes. It differs from C. erythropoda little other than in leaf shape and ripe fruit color. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 6 (6 in the flora). Crataegus pruinosa extends from Arkansas to Wisconsin, through the southern Great Lakes to southern New England, and, in the south, mainly in the Appalachians to northern Georgia. In the north of its range, Crataegus pruinosa is mainly a shrub of open successional habitats but in the south may commonly be a taller tree of open or thin woodlands. Many hawthorns have a little waxy bloom on their pomes; it is particularly prominent on Crataegus pruinosa and C. cognata compared to others. Some authors include in C. pruinosa their white-anthered counterparts, here assigned to C. cognata. Whereas C. pruinosa characteristically has entire or subentire sepal margins, some forms in the southwest of the range of the species (for example, C. calliantha Sargent, C. seducta Sargent) may have glandular-serrate sepal margins. They may represent introgression with C. coccinioides. The varieties of C. pruinosa are weakly differentiated from each other, most of them on leaf shape and size characters. The more widespread varieties constitute a range of morphotypes held together by common traits. Crataegus gaudens Sargent is a strikingly distinct form from Pennsylvania that has more or less elliptic leaves with lobes absent; it is clearly related to C. pruinosa. Note that 159. Crataegus ×coleae, a Michigan endemic, will key out here if its laterally scarred pyrenes are missed. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||||||||||||||||||
Key |
|
|||||||||||||||||||||
Source | FNA vol. 9, p. 510. | FNA vol. 9, p. 574. | ||||||||||||||||||||
Parent taxa | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Douglasia > ser. Cerrones | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Pruinosae | ||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||
Synonyms | Mespilus pruinosa | |||||||||||||||||||||
Name authority | Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 464. (1840) | (H. L. Wendland) K. Koch: Hort. Dendrol., 168. (1853) | ||||||||||||||||||||
Web links |