Flora of North America species comparison

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Crataegus rivularis

Crataegus ashei

mountain river hawthorn, river hawthorn

Ashe hawthorn
Habit Shrubs or trees, 30–50 dm. Shrubs, 30–50 dm.
Stems

twigs: new growth greenish, sparsely pubescent, early glabrescent, 1–2-years old often red-purple;

bark on younger 2–5 cm thick branches dark gray-brown, sometimes copper-colored;

thorns on twigs straight or ± recurved, 2-years old black or purple-black, glossy, fine, 1.5–4 cm.

twigs: new growth densely pubescent, 1-year old tan to chestnut, ± shiny, older gray;

thorns on twigs few to numerous, 2-years old shiny, very dark brown to black, ± slender, 2–3 cm.
Leaves

petiole 1–2.5 cm, glabrous;

blade elliptic to narrowly elliptic, 3–8 cm, length 2+ times width, thin, base cuneate, lobes 0 or small apiculi at ends of some vein tips, margins serrate, teeth numerous, usually acute, fine to coarse, venation semicamptodromous, veins 4 or 5 per side, larger often branched proximal to ends, apex acute, abaxial surface glabrous, adaxial appressed-strigose young.

petiole length 25% blade, densely pubescent, stipitate-glandular;

blade mid green, elliptic to narrowly obovate, 3–6 cm, base cuneate, lobes 0 or obscure, margins glandular-serrate, teeth to 1.5 mm, proximal teeth stipitate-glandular, veins 5–7 per side, apex acute to subacute, abaxial surface sparsely pubescent or glabrous, veins hairy, adaxial shiny, scabrous.
Inflorescences

6–12-flowered;

branches glandular-punctate;

bracteoles early caducous, usually present, margins sessile-glandular.

3–10-flowered, arising proleptically (subterminal on perennial short shoots);

branches densely villous;

bracteoles caducous, linear, 4–8 mm, herbaceous to membranous, margins stipitate-glandular.
Flowers

14–17 mm diam.;

sepals broadly deltate, 6–8 mm, margins entire, apex narrow, glabrous;

stamens 10, anthers pink, sometimes ivory;

styles 3–5.

(15–)20–23 mm diam.;

hypanthium tomentose;

sepals narrowly triangular, 6–7 mm, margins glandular-pectinate, abaxially pilose;

stamens 20(–25), anthers cream, sometimes pink;

styles 3(–5) (with bristly hairs at bases).
Pomes

deep red (mid Aug), black or blackish purple mature, suborbicular, 10 mm diam.;

sepals ± erect to strongly outcurved or ± patent, 2–3 mm;

pyrenes 3 or 4, sides eroded.

orange-red to deep red, 10–14 mm diam., densely short-pubescent;

sepals patent-incurved;

pyrenes 3(–5).
2n

= 68.

Crataegus rivularis

Crataegus ashei
Phenology Flowering mid May–early Jun; fruiting Aug–Sep. Flowering Apr; fruiting Sep–Nov.
Habitat Intermontane streamsides, ditches, flood plains Brush, open woodlands
Elevation 1300–2300 m (4300–7500 ft) 0–300 m (0–1000 ft)
Distribution
from FNA
AZ; CO; ID; NM; NV; UT; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; LA; MS; TN
[BONAP county map]
Discussion

Crataegus rivularis is most abundant in intermontane Colorado and Utah and is one of two hawthorns found in Arizona; it is rare in Nevada. There are reports from northwestern Texas (for example, D. S. Correll and M. C. Johnston 1970); they have not been verified. Crataegus rivularis is one of the more common woody species of mesic, valley bottom intermontane habitats. The species flowers early.

Crataegus rivularis is superficially similar to the more or less sympatric, less common, C. saligna; it has larger leaves with fewer veins, larger flowers with ten stamens, and larger pomes. It differs from C. erythropoda little other than in leaf shape and ripe fruit color.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Crataegus ashei occurs from Louisiana to Alabama and is apparently scarce.

Crataegus ashei has often been confused with both C. triflora (ser. Triflorae) and C. harbisonii; it differs from C. triflora in its different growth habit with branching more or less layered and leaves smallish, flowers smaller, stamens 20–24, and leaves usually more coriaceous and shiny with lobes absent. Crataegus ashei is more similar to C. harbisonii, mainly differing in being smaller in its vegetative parts, invariably lacking reproductive side shoots of the season, and by having entire leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 9, p. 510. FNA vol. 9, p. 608.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Douglasia > ser. Cerrones Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Bracteatae
Sibling taxa
C. aemula, C. aestivalis, C. alabamensis, C. alleghaniensis, C. annosa, C. aprica, C. aquacervensis, C. ashei, C. atrovirens, C. attrita, C. austromontana, C. beata, C. berberifolia, C. biltmoreana, C. brachyacantha, C. brainerdii, C. brazoria, C. brittonii, C. buckleyi, C. calpodendron, C. castlegarensis, C. chrysocarpa, C. coccinea, C. coccinioides, C. cognata, C. collina, C. colonica, C. communis, C. compacta, C. condigna, C. craytonii, C. crocea, C. crus-galli, C. cupressocollina, C. delawarensis, C. dispar, C. dodgei, C. douglasii, C. egens, C. egregia, C. enderbyensis, C. erythropoda, C. exilis, C. eximia, C. extraria, C. fecunda, C. flabellata, C. flava, C. florens, C. floridana, C. florifera, C. fluviatilis, C. formosa, C. frugiferens, C. furtiva, C. gattingeri, C. gaylussacia, C. gilva, C. greggiana, C. harbisonii, C. holmesiana, C. ignava, C. incilis, C. integra, C. intricata, C. invicta, C. iracunda, C. irrasa, C. jesupii, C. jonesiae, C. lacrimata, C. laevigata, C. lanata, C. lancei, C. lanuginosa, C. lassa, C. laurentiana, C. leonensis, C. lepida, C. levis, C. lumaria, C. macracantha, C. macrosperma, C. magniflora, C. margarettae, C. marshallii, C. mendosa, C. meridiana, C. mira, C. mollis, C. monogyna, C. munda, C. nananixonii, C. neobushii, C. nitida, C. oakesiana, C. okanaganensis, C. okennonii, C. opaca, C. opima, C. orbicularis, C. ouachitensis, C. padifolia, C. pennsylvanica, C. persimilis, C. pexa, C. phaenopyrum, C. phippsii, C. pinetorum, C. populnea, C. prona, C. pruinosa, C. pulcherrima, C. punctata, C. purpurella, C. quaesita, C. reverchonii, C. rivuloadamensis, C. rivulopugnensis, C. roribacca, C. rubella, C. rubribracteolata, C. saligna, C. sargentii, C. scabrida, C. schizophylla, C. schuettei, C. segnis, C. senta, C. sheila-phippsiae, C. sheridana, C. shuswapensis, C. sororia, C. spathulata, C. spes-aestatum, C. stolonifera, C. stonei, C. submollis, C. suborbiculata, C. succulenta, C. tecta, C. teres, C. texana, C. tracyi, C. triflora, C. turnerorum, C. uniflora, C. ursopedensis, C. venusta, C. viridis, C. visenda, C. wattiana, C. williamsii, C. wootoniana, C. ×atrorubens, C. ×bicknellii, C. ×coleae, C. ×collicola, C. ×disperma, C. ×dispessa, C. ×fretalis, C. ×incaedua, C. ×kelloggii, C. ×latebrosa, C. ×lucorum, C. ×rufula, C. ×sicca, C. ×vailiae
C. aemula, C. aestivalis, C. alabamensis, C. alleghaniensis, C. annosa, C. aprica, C. aquacervensis, C. atrovirens, C. attrita, C. austromontana, C. beata, C. berberifolia, C. biltmoreana, C. brachyacantha, C. brainerdii, C. brazoria, C. brittonii, C. buckleyi, C. calpodendron, C. castlegarensis, C. chrysocarpa, C. coccinea, C. coccinioides, C. cognata, C. collina, C. colonica, C. communis, C. compacta, C. condigna, C. craytonii, C. crocea, C. crus-galli, C. cupressocollina, C. delawarensis, C. dispar, C. dodgei, C. douglasii, C. egens, C. egregia, C. enderbyensis, C. erythropoda, C. exilis, C. eximia, C. extraria, C. fecunda, C. flabellata, C. flava, C. florens, C. floridana, C. florifera, C. fluviatilis, C. formosa, C. frugiferens, C. furtiva, C. gattingeri, C. gaylussacia, C. gilva, C. greggiana, C. harbisonii, C. holmesiana, C. ignava, C. incilis, C. integra, C. intricata, C. invicta, C. iracunda, C. irrasa, C. jesupii, C. jonesiae, C. lacrimata, C. laevigata, C. lanata, C. lancei, C. lanuginosa, C. lassa, C. laurentiana, C. leonensis, C. lepida, C. levis, C. lumaria, C. macracantha, C. macrosperma, C. magniflora, C. margarettae, C. marshallii, C. mendosa, C. meridiana, C. mira, C. mollis, C. monogyna, C. munda, C. nananixonii, C. neobushii, C. nitida, C. oakesiana, C. okanaganensis, C. okennonii, C. opaca, C. opima, C. orbicularis, C. ouachitensis, C. padifolia, C. pennsylvanica, C. persimilis, C. pexa, C. phaenopyrum, C. phippsii, C. pinetorum, C. populnea, C. prona, C. pruinosa, C. pulcherrima, C. punctata, C. purpurella, C. quaesita, C. reverchonii, C. rivularis, C. rivuloadamensis, C. rivulopugnensis, C. roribacca, C. rubella, C. rubribracteolata, C. saligna, C. sargentii, C. scabrida, C. schizophylla, C. schuettei, C. segnis, C. senta, C. sheila-phippsiae, C. sheridana, C. shuswapensis, C. sororia, C. spathulata, C. spes-aestatum, C. stolonifera, C. stonei, C. submollis, C. suborbiculata, C. succulenta, C. tecta, C. teres, C. texana, C. tracyi, C. triflora, C. turnerorum, C. uniflora, C. ursopedensis, C. venusta, C. viridis, C. visenda, C. wattiana, C. williamsii, C. wootoniana, C. ×atrorubens, C. ×bicknellii, C. ×coleae, C. ×collicola, C. ×disperma, C. ×dispessa, C. ×fretalis, C. ×incaedua, C. ×kelloggii, C. ×latebrosa, C. ×lucorum, C. ×rufula, C. ×sicca, C. ×vailiae
Name authority Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 464. (1840) Beadle: Bot. Gaz. 30: 339. (1900)
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Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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