Crataegus pruinosa |
Crataegus ×disperma |
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frosted hawthorn, waxyfruit hawthorn |
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Habit | Shrubs or trees, dense, often suberect, 20–70 dm. | Shrubs or trees, 50–80 dm. | ||||||||||||||||||||
Stems | compound thorns on trunks present; twigs: new growth reddish brown, 1-year old dull purple-brown, 2-years old dull gray, older paler; thorns on twigs straight to slightly recurved, 2-years old deep purple to shiny black, usually ± fine, 3–5 cm. |
twigs: new growth reddish young, glabrous or pubescent, 1-year old shiny brown, glabrous, older dark gray; thorns on twigs 2-years old shiny, dark brown, fine, 3–4 cm. |
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Leaves | petiole length 50–66% blade, sparsely glandular; blade trullate to ovate, ovate-oblong, or broadly rhombic, rarely ± deltate, 2–6(–7) cm, subcoriaceous, base broadly cuneate to subtruncate to weakly subcordate, lobes (1–)3 or 4 per side, sinuses shallow to deeper, lobe apex acute, sometimes ± obtuse, margins serrate, veins 5 or 6 per side (except smaller leaves), apex acute, surfaces glabrous (except var. virella). |
deciduous; petiole 0.7–1.5 cm, winged nearly to base, glabrous; blade dark green, obovate or elliptic, 2.5–5 cm, subcoriaceous, base broadly to narrowly cuneate, lobes 0 or few, sinuses very shallow, margins serrate, venation craspedodromous, veins 5–7 per side, apex acute, surfaces glabrous. |
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Inflorescences | 5–10-flowered; branches glabrous, sometimes sparsely villous; bracteoles caducous, usually few, margins short-stipitate-glandular. |
5–12-flowered; branches glabrous; bracteoles caducous, linear, membranous, margins glandular. |
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Flowers | 15–25 mm diam.; sepals narrowly triangular, 5–6 mm, margins usually entire or subentire, rarely glandular-serrate, abaxially glabrous; stamens (10 or)20, anthers pale pink to bright rose or dull purple, sometimes cream, 0.6–0.8 mm; styles 3–5. |
13–18 mm diam.; hypanthium glabrous; sepals narrow, margins ± entire or subserrate; stamens 10, anthers pink or red-purple; styles 2(or 3). |
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Pomes | greenish with pink or mauve areas, sometimes bright crimson or scarlet, often rather angular, 10–20 mm diam., highly pruinose, not punctate; flesh hard; sepals on collar, spreading; pyrenes 3–5. |
deep red, ± oblong, 12–14 × 9–11 mm; sepals spreading, subsessile; pyrenes 2, dorsally grooved, sides plane. |
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Crataegus pruinosa |
Crataegus ×disperma |
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Phenology | Flowering May; fruiting Sep–Oct. | |||||||||||||||||||||
Habitat | Brush, along streams | |||||||||||||||||||||
Elevation | 50–200 m (200–700 ft) | |||||||||||||||||||||
Distribution |
AR; CT; GA; IA; IL; IN; KS; KY; MA; ME; MI; MO; NC; NH; NJ; NY; OH; PA; TN; VA; VT; WI; WV; ON; QC
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IA; IL; IN; OH; PA; VA |
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Discussion | Varieties 6 (6 in the flora). Crataegus pruinosa extends from Arkansas to Wisconsin, through the southern Great Lakes to southern New England, and, in the south, mainly in the Appalachians to northern Georgia. In the north of its range, Crataegus pruinosa is mainly a shrub of open successional habitats but in the south may commonly be a taller tree of open or thin woodlands. Many hawthorns have a little waxy bloom on their pomes; it is particularly prominent on Crataegus pruinosa and C. cognata compared to others. Some authors include in C. pruinosa their white-anthered counterparts, here assigned to C. cognata. Whereas C. pruinosa characteristically has entire or subentire sepal margins, some forms in the southwest of the range of the species (for example, C. calliantha Sargent, C. seducta Sargent) may have glandular-serrate sepal margins. They may represent introgression with C. coccinioides. The varieties of C. pruinosa are weakly differentiated from each other, most of them on leaf shape and size characters. The more widespread varieties constitute a range of morphotypes held together by common traits. Crataegus gaudens Sargent is a strikingly distinct form from Pennsylvania that has more or less elliptic leaves with lobes absent; it is clearly related to C. pruinosa. Note that 159. Crataegus ×coleae, a Michigan endemic, will key out here if its laterally scarred pyrenes are missed. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Crataegus ×disperma has been found sporadically from Iowa east to Virginia and Pennsylvania, where the ranges of C. punctata and C. crus-galli overlap. Crataegus ×disperma belongs to a group of taxa intermediate between C. crus-galli and C. punctata; it may key to C. collina, although it has glossier leaves and usually fewer styles and pyrenes. Some specimens are more hairy, or with more strongly glandular-serrate sepal margins, than in the description, which is based on Ashe. Crataegus danielsii E. J. Palmer from Missouri, with glossy, more or less coriaceous, but lobed leaves is another problematic C. crus-galli × C. collina hybrid according to Palmer (1952). The pink-anthered C. ×disperma and cream-anthered C. ×incaedua are similar to C. collina var. sordida except for the number of styles and pyrenes (two or three). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 574. | FNA vol. 9, p. 637. | ||||||||||||||||||||
Parent taxa | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Pruinosae | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus | ||||||||||||||||||||
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Synonyms | Mespilus pruinosa | C. grandis, C. pausiaca, C. peoriensis, C. punctata var. pausiaca | ||||||||||||||||||||
Name authority | (H. L. Wendland) K. Koch: Hort. Dendrol., 168. (1853) | Crataegus ×disperma Ashe: J. Elisha Mitchell Sci. Soc. 17: 14. (1900) — as species | ||||||||||||||||||||
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