frosted hawthorn, waxyfruit hawthorn
|
Wooton's hawthorn
|
Shrubs or trees, dense, often suberect, 20–70 dm. |
Shrubs, 20 dm. |
compound thorns on trunks present; twigs: new growth reddish brown, 1-year old dull purple-brown, 2-years old dull gray, older paler; thorns on twigs straight to slightly recurved, 2-years old deep purple to shiny black, usually ± fine, 3–5 cm. |
little branched, thorny; twigs: new growth not recorded, glabrous, 1-year old reddish brown, older gray; thorns on twigs 1-year old reddish brown, ± stout, 4 cm. |
petiole length 50–66% blade, sparsely glandular; blade trullate to ovate, ovate-oblong, or broadly rhombic, rarely ± deltate, 2–6(–7) cm, subcoriaceous, base broadly cuneate to subtruncate to weakly subcordate, lobes (1–)3 or 4 per side, sinuses shallow to deeper, lobe apex acute, sometimes ± obtuse, margins serrate, veins 5 or 6 per side (except smaller leaves), apex acute, surfaces glabrous (except var. virella). |
petiole length 40–50% blade, eglandular or sparsely glandular; blade ovate, 3–4(–5) cm, 70–100% mature size at anthesis, base truncate to broadly cuneate, lobes 3 or 4 per side, sinuses shallow, lobe apex obtuse to cuspidate, margins serrate, veins 4 or 5 per side, apex subacute, adaxial surface short-pubescent young, later glabrate. |
5–10-flowered; branches glabrous, sometimes sparsely villous; bracteoles caducous, usually few, margins short-stipitate-glandular. |
5–10-flowered; branches glabrous; bracteoles absent or few, shape not recorded; anthesis in spring. |
15–25 mm diam.; sepals narrowly triangular, 5–6 mm, margins usually entire or subentire, rarely glandular-serrate, abaxially glabrous; stamens (10 or)20, anthers pale pink to bright rose or dull purple, sometimes cream, 0.6–0.8 mm; styles 3–5. |
10–12 mm diam.; hypanthium glabrous; sepals 2–3 mm, margins subentire, abaxially glabrous; stamens 10, anthers pink; styles 3. |
greenish with pink or mauve areas, sometimes bright crimson or scarlet, often rather angular, 10–20 mm diam., highly pruinose, not punctate; flesh hard; sepals on collar, spreading; pyrenes 3–5. |
shiny red, ellipsoid, 8 mm; sepal remnants erect-patent; pyrenes 2 or 3, dorsally grooved. |
|
|
|
|
|
Flowering May; fruiting Sep. |
|
Grassy areas near streams |
|
2000–3000 m (6600–9800 ft) |
AR; CT; GA; IA; IL; IN; KS; KY; MA; ME; MI; MO; NC; NH; NJ; NY; OH; PA; TN; VA; VT; WI; WV; ON; QC
|
NM |
Varieties 6 (6 in the flora). Crataegus pruinosa extends from Arkansas to Wisconsin, through the southern Great Lakes to southern New England, and, in the south, mainly in the Appalachians to northern Georgia. In the north of its range, Crataegus pruinosa is mainly a shrub of open successional habitats but in the south may commonly be a taller tree of open or thin woodlands. Many hawthorns have a little waxy bloom on their pomes; it is particularly prominent on Crataegus pruinosa and C. cognata compared to others. Some authors include in C. pruinosa their white-anthered counterparts, here assigned to C. cognata. Whereas C. pruinosa characteristically has entire or subentire sepal margins, some forms in the southwest of the range of the species (for example, C. calliantha Sargent, C. seducta Sargent) may have glandular-serrate sepal margins. They may represent introgression with C. coccinioides. The varieties of C. pruinosa are weakly differentiated from each other, most of them on leaf shape and size characters. The more widespread varieties constitute a range of morphotypes held together by common traits. Crataegus gaudens Sargent is a strikingly distinct form from Pennsylvania that has more or less elliptic leaves with lobes absent; it is clearly related to C. pruinosa. Note that 159. Crataegus ×coleae, a Michigan endemic, will key out here if its laterally scarred pyrenes are missed. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Crataegus wootoniana is known from New Mexico. No authentic records of C. wootoniana for Arizona are known; all alleged examples seen proved to be C. macracantha. Crataegus wootoniana is similar to the widespread 57. C. macrosperma (see discussion thereunder), from which it differs little. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| var. dissona |
| → 2 |
2. Leaf surfaces sparsely short-hairy adaxially young, usually glabrescent; inflorescence branches sometimes sparsely villous. | var. virella |
2. Leaf surfaces glabrous adaxially young, sometimes sparsely appressed-hairy along veins; inflorescence branches glabrous | → 3 |
3. Leaf blades 5–7 cm, ovate to ovate-deltate, lobes 0 or 1–4 per side, obscure, lobe apices ± obtuse, bases broadly cuneate to subtruncate; anthers cream. | var. magnifolia |
3. Leaf blades 2–7 cm, ovate or ovate-oblong to ± deltate or broadly rhombic, lobes 2–4 per side, distinct, lobe apices acute, bases broadly cuneate to subtruncate or truncate to weakly subcordate; anthers pale pink | → 4 |
4. Leaf blades 3–7 cm, broadly ovate to deltate, length/width = 1–1.2, bases very broadly cuneate or truncate to weakly subcordate. | var. rugosa |
4. Leaf blades 2–6 cm, broadly rhombic, ovate, or ovate-oblong to narrowly deltate, length/width = 1.5, bases broadly cuneate to subtruncate | → 5 |
5. Leaf blades ovate or ovate-oblong to narrowly deltate, 4–6 cm, bases broadly cuneate to subtruncate. | var. pruinosa |
5. Leaf blades broadly rhombic, 2–4 cm, bases broadly cuneate. | var. parvula |
|
|
FNA vol. 9, p. 574. |
FNA vol. 9, p. 564. |
Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Pruinosae |
Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Tenuifoliae |
C. aemula, C. aestivalis, C. alabamensis, C. alleghaniensis, C. annosa, C. aprica, C. aquacervensis, C. ashei, C. atrovirens, C. attrita, C. austromontana, C. beata, C. berberifolia, C. biltmoreana, C. brachyacantha, C. brainerdii, C. brazoria, C. brittonii, C. buckleyi, C. calpodendron, C. castlegarensis, C. chrysocarpa, C. coccinea, C. coccinioides, C. cognata, C. collina, C. colonica, C. communis, C. compacta, C. condigna, C. craytonii, C. crocea, C. crus-galli, C. cupressocollina, C. delawarensis, C. dispar, C. dodgei, C. douglasii, C. egens, C. egregia, C. enderbyensis, C. erythropoda, C. exilis, C. eximia, C. extraria, C. fecunda, C. flabellata, C. flava, C. florens, C. floridana, C. florifera, C. fluviatilis, C. formosa, C. frugiferens, C. furtiva, C. gattingeri, C. gaylussacia, C. gilva, C. greggiana, C. harbisonii, C. holmesiana, C. ignava, C. incilis, C. integra, C. intricata, C. invicta, C. iracunda, C. irrasa, C. jesupii, C. jonesiae, C. lacrimata, C. laevigata, C. lanata, C. lancei, C. lanuginosa, C. lassa, C. laurentiana, C. leonensis, C. lepida, C. levis, C. lumaria, C. macracantha, C. macrosperma, C. magniflora, C. margarettae, C. marshallii, C. mendosa, C. meridiana, C. mira, C. mollis, C. monogyna, C. munda, C. nananixonii, C. neobushii, C. nitida, C. oakesiana, C. okanaganensis, C. okennonii, C. opaca, C. opima, C. orbicularis, C. ouachitensis, C. padifolia, C. pennsylvanica, C. persimilis, C. pexa, C. phaenopyrum, C. phippsii, C. pinetorum, C. populnea, C. prona, C. pulcherrima, C. punctata, C. purpurella, C. quaesita, C. reverchonii, C. rivularis, C. rivuloadamensis, C. rivulopugnensis, C. roribacca, C. rubella, C. rubribracteolata, C. saligna, C. sargentii, C. scabrida, C. schizophylla, C. schuettei, C. segnis, C. senta, C. sheila-phippsiae, C. sheridana, C. shuswapensis, C. sororia, C. spathulata, C. spes-aestatum, C. stolonifera, C. stonei, C. submollis, C. suborbiculata, C. succulenta, C. tecta, C. teres, C. texana, C. tracyi, C. triflora, C. turnerorum, C. uniflora, C. ursopedensis, C. venusta, C. viridis, C. visenda, C. wattiana, C. williamsii, C. wootoniana, C. ×atrorubens, C. ×bicknellii, C. ×coleae, C. ×collicola, C. ×disperma, C. ×dispessa, C. ×fretalis, C. ×incaedua, C. ×kelloggii, C. ×latebrosa, C. ×lucorum, C. ×rufula, C. ×sicca, C. ×vailiae |
C. aemula, C. aestivalis, C. alabamensis, C. alleghaniensis, C. annosa, C. aprica, C. aquacervensis, C. ashei, C. atrovirens, C. attrita, C. austromontana, C. beata, C. berberifolia, C. biltmoreana, C. brachyacantha, C. brainerdii, C. brazoria, C. brittonii, C. buckleyi, C. calpodendron, C. castlegarensis, C. chrysocarpa, C. coccinea, C. coccinioides, C. cognata, C. collina, C. colonica, C. communis, C. compacta, C. condigna, C. craytonii, C. crocea, C. crus-galli, C. cupressocollina, C. delawarensis, C. dispar, C. dodgei, C. douglasii, C. egens, C. egregia, C. enderbyensis, C. erythropoda, C. exilis, C. eximia, C. extraria, C. fecunda, C. flabellata, C. flava, C. florens, C. floridana, C. florifera, C. fluviatilis, C. formosa, C. frugiferens, C. furtiva, C. gattingeri, C. gaylussacia, C. gilva, C. greggiana, C. harbisonii, C. holmesiana, C. ignava, C. incilis, C. integra, C. intricata, C. invicta, C. iracunda, C. irrasa, C. jesupii, C. jonesiae, C. lacrimata, C. laevigata, C. lanata, C. lancei, C. lanuginosa, C. lassa, C. laurentiana, C. leonensis, C. lepida, C. levis, C. lumaria, C. macracantha, C. macrosperma, C. magniflora, C. margarettae, C. marshallii, C. mendosa, C. meridiana, C. mira, C. mollis, C. monogyna, C. munda, C. nananixonii, C. neobushii, C. nitida, C. oakesiana, C. okanaganensis, C. okennonii, C. opaca, C. opima, C. orbicularis, C. ouachitensis, C. padifolia, C. pennsylvanica, C. persimilis, C. pexa, C. phaenopyrum, C. phippsii, C. pinetorum, C. populnea, C. prona, C. pruinosa, C. pulcherrima, C. punctata, C. purpurella, C. quaesita, C. reverchonii, C. rivularis, C. rivuloadamensis, C. rivulopugnensis, C. roribacca, C. rubella, C. rubribracteolata, C. saligna, C. sargentii, C. scabrida, C. schizophylla, C. schuettei, C. segnis, C. senta, C. sheila-phippsiae, C. sheridana, C. shuswapensis, C. sororia, C. spathulata, C. spes-aestatum, C. stolonifera, C. stonei, C. submollis, C. suborbiculata, C. succulenta, C. tecta, C. teres, C. texana, C. tracyi, C. triflora, C. turnerorum, C. uniflora, C. ursopedensis, C. venusta, C. viridis, C. visenda, C. wattiana, C. williamsii, C. ×atrorubens, C. ×bicknellii, C. ×coleae, C. ×collicola, C. ×disperma, C. ×dispessa, C. ×fretalis, C. ×incaedua, C. ×kelloggii, C. ×latebrosa, C. ×lucorum, C. ×rufula, C. ×sicca, C. ×vailiae |
|
|
Mespilus pruinosa |
|
(H. L. Wendland) K. Koch: Hort. Dendrol., 168. (1853) |
Eggleston: Torreya 7: 236. (1908) |
| |