Crataegus holmesiana |
Crataegus rivuloadamensis |
|
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aubépine de Holmes, Holmes' hawthorn |
Adams Creek hawthorn |
|
Habit | Shrubs or trees, 40–70(–100) dm, single-trunked. | Shrubs or trees, 25–40(–50) dm, wide-branching. |
Stems | twigs: new growth greenish to reddish, glabrous or pubescent; thorns on twigs recurved, often slender, 3–6 cm. |
twigs: new growth dark reddish green, pubescent, 1-year old shiny, dark red-brown, 2-years old dark reddish and gray; thorns on twigs ± straight, 1-year old glossy, very dark red-brown, older dark gray, shorter conic, stout, 1.5–2.5(–3) cm. |
Leaves | petiole length 30–35% blade, glabrate to densely hairy, often sparsely glandular; blade ± narrowly ovate, 6–9 cm, base cuneate or ± rounded, lobe apex acuminate, margins serrate or doubly serrate, teeth 2 mm, adaxial usually densely scabrous-pubescent young. |
petiole length 25–35% blade, eglandular or sparsely sessile-glandular, stipitate-glandular at leaf base, adaxial sulcus densely pubescent; blade ± elliptic or broadly elliptic to rhombic-elliptic, 2–4 cm at anthesis, 4–8 cm mature, base cuneate to narrowly cuneate, lobes 2–4 per side, max LII (0–)5–20(–25)%, lobe apex subacute to broadly cuspidate, margins finely crenate-serrate or serrate, veins 4–6 per side, hardly impressed, apex acute, abaxial surface glabrous or sparsely hairy young, veins appressed-scabrous, adaxial ± densely appressed-scabrous (indumentum persisting). |
Inflorescences | branches usually pubescent, rarely glabrous or tomentose. |
4–10-flowered; branches ± densely tomentose; bracteoles ± reddish, narrow, margins densely sessile-glandular. |
Flowers | hypanthium usually pubescent; stamens 5–8(–10), anthers pink to rose-purple, sometimes crimson. |
17–21 mm diam.; hypanthium densely pubescent except sometimes distally; sepal color not recorded, broadly triangular, 3 mm, margins greenish, abaxial surface glabrous, adaxial pubescent; petals pale cream; anthers pink; styles 3–5. |
Pomes | bright to deep red, ellipsoid to broadly pyriform, 12–14 mm, usually glabrous; sepals wide-spreading or erose. |
1–5(–6) per infructescence, pale reddish purple, becoming deep red to reddish plum and burgundy, suborbicular to broadly turbinate, 10–12 mm, usually ± densely hairy; sepals ± patent to recurved, 4–5 mm; pyrenes 3–5, dorsally strongly grooved, sides plane to very shallowly concave. |
2n | = 34, 51. |
|
Crataegus holmesiana |
Crataegus rivuloadamensis |
|
Phenology | Flowering May; fruiting Sep–Oct. | Flowering late May–early Jun; fruiting Aug–Sep. |
Habitat | Woodland edges, old pastures, fencerows | Thickets, under light shade of aspen |
Elevation | 10–300 m (0–1000 ft) | 800–1200 m (2600–3900 ft) |
Distribution |
CT; IL; IN; MA; ME; MI; MN; NH; NY; OH; PA; RI; VT; WI; ON; QC
|
AB; SK |
Discussion | Crataegus holmesiana ranges from Wisconsin and Illinois through the southern Great Lakes region to southern Quebec and Maine, and to Pennsylvania; it is common in much of its range and should be expected in West Virginia. In its usual form, Crataegus holmesiana is readily distinguished from C. coccinea vars. coccinea and pringlei, though occasional intermediates occur. Crataegus robesoniana Sargent [C. pedicellata var. robesoniana (Sargent) E. J. Palmer] is one and perhaps C. amicta Ashe [C. holmesiana var. amicta (Ashe) E. J. Palmer] is another. Crataegus holmesiana is routinely interpreted as a typical member of ser. Coccineae, thus, a villous inflorescence taxon with serrate sepals. However, the protologue by Ashe (authentic material not located) states that the inflorescences are glabrous and the sepals are nearly entire, causing confusion with C. tenuifolia Britton (ser. Tenuifoliae). Current usage for C. holmesiana became fixed after Ashe erected a var. villosa with serrate sepals and villous inflorescences. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Crataegus rivuloadamensis is known only from the western half of the Cypress Hills and is most abundant at elevations above 1150 m. The extremely dark young wood and short and conic thorns show strong resemblance to species of the Eurasian sect. Sanguineae Zabel ex C. K. Schneider. The unpitted pyrenes are a marked difference. The petals appear to be pale cream rather than pure white. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 560. | FNA vol. 9, p. 634. |
Parent taxa | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Coccineae | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Unassigned > ser. Montaninsulae |
Sibling taxa | ||
Synonyms | C. anomala, C. eamesii, C. elongata, C. holmesiana var. villipes, C. pura, C. sertata, C. tardipes | |
Name authority | Ashe: J. Elisha Mitchell Sci. Soc. 16: 78. (1900) | J. B. Phipps & O’Kennon: J. Bot. Res. Inst. Texas 1: 1065, plate 7.3a, fig. 13. (2007) |
Web links |