Crataegus holmesiana |
Crataegus jonesiae |
|
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aubépine de Holmes, Holmes' hawthorn |
Jones hawthorn, Miss Jones' hawthorn |
|
Habit | Shrubs or trees, 40–70(–100) dm, single-trunked. | Shrubs or trees, 60–70 dm. |
Stems | twigs: new growth greenish to reddish, glabrous or pubescent; thorns on twigs recurved, often slender, 3–6 cm. |
twigs: new growth ± densely appressed-pubescent, 1-year old pale tan, 2-years old olive gray, older gray; thorns on twigs: frequency not recorded, straight to recurved, 1-year old chestnut brown, 2-years old shiny dark brown or black, slender to stout, 3–6 cm. |
Leaves | petiole length 30–35% blade, glabrate to densely hairy, often sparsely glandular; blade ± narrowly ovate, 6–9 cm, base cuneate or ± rounded, lobe apex acuminate, margins serrate or doubly serrate, teeth 2 mm, adaxial usually densely scabrous-pubescent young. |
petiole 1.5–3.5 cm, length 35–50% blade, pubescent, eglandular; blade broadly ovate to broadly rhombic-elliptic or nearly circular, 4–7(–8) cm, length/width = 1–1.5, base tapered, wedge-shaped to subtruncate, lobes (4 or)5–9 per side, obscure, lobe apex usually acute, margins doubly serrate, veins 5–10 per side, apex acute, abaxial surface densely appressed-pubescent on veins, adaxial appressed-pubescent young, sparsely pubescent or glabrescent later. |
Inflorescences | branches usually pubescent, rarely glabrous or tomentose. |
8–15(–20)-flowered, lax; branches with antrorse pubescence; bracteoles membranous, margins glandular. |
Flowers | hypanthium usually pubescent; stamens 5–8(–10), anthers pink to rose-purple, sometimes crimson. |
18–23 mm diam.; hypanthium densely pubescent; sepals 6–8 mm, margins ± entire, abaxially pubescent; stamens 10, anthers pink; styles 2 or 3(or 4). |
Pomes | bright to deep red, ellipsoid to broadly pyriform, 12–14 mm, usually glabrous; sepals wide-spreading or erose. |
bright red, oblong or obovoid, 10–15 mm diam., pubescent, glabrescent; sepals erect-patent; pyrenes 2 or 3(or 4). |
2n | = 34, 51. |
|
Crataegus holmesiana |
Crataegus jonesiae |
|
Phenology | Flowering May; fruiting Sep–Oct. | Flowering Jun; fruiting Sep–Oct. |
Habitat | Woodland edges, old pastures, fencerows | Woodland glades, stream borders, rocky banks |
Elevation | 10–300 m (0–1000 ft) | 10–200 m (0–700 ft) |
Distribution |
CT; IL; IN; MA; ME; MI; MN; NH; NY; OH; PA; RI; VT; WI; ON; QC
|
ME; NB; NS; PE |
Discussion | Crataegus holmesiana ranges from Wisconsin and Illinois through the southern Great Lakes region to southern Quebec and Maine, and to Pennsylvania; it is common in much of its range and should be expected in West Virginia. In its usual form, Crataegus holmesiana is readily distinguished from C. coccinea vars. coccinea and pringlei, though occasional intermediates occur. Crataegus robesoniana Sargent [C. pedicellata var. robesoniana (Sargent) E. J. Palmer] is one and perhaps C. amicta Ashe [C. holmesiana var. amicta (Ashe) E. J. Palmer] is another. Crataegus holmesiana is routinely interpreted as a typical member of ser. Coccineae, thus, a villous inflorescence taxon with serrate sepals. However, the protologue by Ashe (authentic material not located) states that the inflorescences are glabrous and the sepals are nearly entire, causing confusion with C. tenuifolia Britton (ser. Tenuifoliae). Current usage for C. holmesiana became fixed after Ashe erected a var. villosa with serrate sepals and villous inflorescences. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Reports of Crataegus jonesiae from Quebec have not been substantiated; vegetatively similar plants from Matapédia and Matane counties have shallowly, laterally scarred pyrenes; sepals are gland-margined and are barely denticulate. Crataegus jonesiae is usually placed in ser. Anomalae. However, it is a particularly distinct species that has smooth pyrenes and other similarities to C. punctata, and the possibility of a hybrid origin should be considered; its leaves range from nearly entire, prototypically punctate, to sharply lobed as in C. chrysocarpa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 560. | FNA vol. 9, p. 545. |
Parent taxa | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Coccineae | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Punctatae |
Sibling taxa | ||
Synonyms | C. anomala, C. eamesii, C. elongata, C. holmesiana var. villipes, C. pura, C. sertata, C. tardipes | |
Name authority | Ashe: J. Elisha Mitchell Sci. Soc. 16: 78. (1900) | Sargent: Bot. Gaz. 31: 14. (1901) |
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