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aubépine flabelliforme, fan-leaf hawthorn
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hoary hawthorn
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| Shrubs or trees, 40–50 dm. |
Shrubs, widely spreading mature, 40–60 dm, branches ± weeping. |
twigs: new growth reddish at first, usually sparsely pubescent, 1-year old deep chestnut brown, older dull gray; thorns on twigs 1-year old shiny, dark brown, stout to slender, 4–6 cm. |
twigs: new growth densely appressed-white-pubescent, 1-year old brown to purple-brown, glabrous; thorns on twigs absent or numerous, straight, 1-year old purple-brown, slender, 1–4 cm. |
petiole 1–2 mm wide, length 40–50% blade, glandular; blade glossy, dark green, broadly elliptic to ovate, 4–7 cm, base cuneate to slightly rounded, lobes 4 or 5 per side, sinuses deep, lobe apex acuminate, margins sharply serrate, teeth gland-tipped, veins 5 or 6 per side, apex acute, abaxial surface glabrous except along veins, adaxial densely appressed-scabrous young, glabrescent. |
petiole length 20–25% blade, pubescent young, densely glandular; blade usually narrowly to broadly obtrullate, sometimes ± obovate, 3–5 cm, thick, floppy, base usually gradually tapered, lobes 1 or 2(or 3) per side, subterminal, short, well defined, margins subentire or obscurely glandular crenate-serrate, veins 2 or 3 per side (exiting beyond widest part of leaf), apex acute to subacute, surfaces pubescent. |
5–10-flowered; branches villous; bracteole frequency not recorded, linear. |
1–4-flowered; branches densely white-canescent; bracteoles deciduous, linear, margins sessile-glandular, adaxially short-pubescent. |
13–17 mm diam.; hypanthium glabrous or villous at base; sepals 4–5 mm, margins entire, glandular to glandular-serrate, rarely nearly eglandular, apex acute, abaxial pubescence not recorded; stamens 5–10 or 20, anthers usually pink; styles 3–5. |
17–20 mm diam.; hypanthium densely canescent; sepals narrowly triangular, 5 mm, margins glandular-serrate, abaxially pubescent; anthers ivory; styles 4 or 5. |
crimson, ± ellipsoid, 8–10 mm diam., sometimes ± pruinose; sepals spreading or usually erose, not or obscurely elevated; pyrenes 3–5, dorsally deeply grooved. |
usually orange or ruddy to crimson, suborbicular to ± pyriform, 9 mm diam., pubescent; sepals reflexed; pyrenes 4 or 5. |
= 51. |
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| Flowering May–Jun; fruiting Sep–Oct. |
Flowering Mar–Apr; fruiting Jul–Aug. |
| Brush, successional fields, fencelines, woodland edges, open woodlands |
Pine barrens, open scrub, on sand |
| 10–200 m (0–700 ft) |
0–200 m (0–700 ft) |
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CT; MA; ME; NH; NY; VT; NB; NS; ON; QC
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AL; FL; GA; NC; SC; VA |
Crataegus flabellata occurs along the Niagara Escarpment in Ontario, on both sides of Lake Ontario and the St. Lawrence, to New Brunswick and Nova Scotia, and to Massachusetts. Reports for Newfoundland have not been substantiated. Crataegus flabellata is distinctive with its leaf blades glossy dark green, ovate to broadly elliptic, cuneate at the base, and with deep acuminate lobes. Specimens with less glandularity or serration of the sepal margins may resemble C. fluviatilis. In spite of the obvious differences in morphology and range, C. flabellata and C. macrosperma often have been combined by floristicians. From C. chrysocarpa, C. flabellata is distinguished by its shinier, ellipsoid, glabrous fruit, by the usually deeper, acuminate lobes of its glossy, dark green leaves, and by longer, slightly more acute sepals. Typically, the inflorescences and hypanthia of eastern C. chrysocarpa are densely hairy, practically tomentose, unlike C. flabellata with its more sparsely hairy pedicels and glabrous distal hypanthia. It is conceivable that, due to morphological intermediacy and the sympatry of all three species within the range of C. flabellata, this species represents the result of ancient hybridization between C. chrysocarpa and C. macrosperma. Crataegus flabellata is usually divided into two varieties: var. flabellata (10 stamens) and var. grayana (20 stamens), the latter being particularly common in southern Quebec. There appears to be little, if any, other constant differentiating feature. Crataegus crudelis, also from Quebec, is a form recognized by long thorns (to 10 cm). The name Crataegus densiflora Sargent, which pertains to C. flabellata, is illegitimate. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Crataegus lanata occurs from Alabama and northern Florida to the Carolinas; there is one record from Chesterfield County, Virginia. Crataegus lanata is one of the most abundant species in ser. Lacrimatae; it is most like C. lassa and is noted for its very large (for ser. Lacrimatae), usually floppy, distinctly but shallowly incised, kite-shaped leaves, persistently somewhat white-hairy. The fruit of C. lanata ripens copper to red. Variants of Crataegus lanata include some intermediates with the typical form of C. lassa and forms with wider, more obovate leaves, some of which may be the same as C. amica Beadle. Crataegus pulla Beadle, described from sandy flats along the Tombigbee River, Mississippi, is a form little-known apart from its type and protologue. Crataegus pulla has similarities to C. lanata, but differs by its more or less coarsely and rather irregularly toothed, obovate leaves often with a single, relatively small, somewhat obscure lobe on each side; the leaves are at first more or less tomentose and then variably somewhat glabrescent; flowers are smaller (14–17 mm wide); and fruits are larger (9–12 mm wide). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 9, p. 567. |
FNA vol. 9, p. 630. |
| Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Tenuifoliae |
Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Lacrimatae |
C. aemula, C. aestivalis, C. alabamensis, C. alleghaniensis, C. annosa, C. aprica, C. aquacervensis, C. ashei, C. atrovirens, C. attrita, C. austromontana, C. beata, C. berberifolia, C. biltmoreana, C. brachyacantha, C. brainerdii, C. brazoria, C. brittonii, C. buckleyi, C. calpodendron, C. castlegarensis, C. chrysocarpa, C. coccinea, C. coccinioides, C. cognata, C. collina, C. colonica, C. communis, C. compacta, C. condigna, C. craytonii, C. crocea, C. crus-galli, C. cupressocollina, C. delawarensis, C. dispar, C. dodgei, C. douglasii, C. egens, C. egregia, C. enderbyensis, C. erythropoda, C. exilis, C. eximia, C. extraria, C. fecunda, C. flava, C. florens, C. floridana, C. florifera, C. fluviatilis, C. formosa, C. frugiferens, C. furtiva, C. gattingeri, C. gaylussacia, C. gilva, C. greggiana, C. harbisonii, C. holmesiana, C. ignava, C. incilis, C. integra, C. intricata, C. invicta, C. iracunda, C. irrasa, C. jesupii, C. jonesiae, C. lacrimata, C. laevigata, C. lanata, C. lancei, C. lanuginosa, C. lassa, C. laurentiana, C. leonensis, C. lepida, C. levis, C. lumaria, C. macracantha, C. macrosperma, C. magniflora, C. margarettae, C. marshallii, C. mendosa, C. meridiana, C. mira, C. mollis, C. monogyna, C. munda, C. nananixonii, C. neobushii, C. nitida, C. oakesiana, C. okanaganensis, C. okennonii, C. opaca, C. opima, C. orbicularis, C. ouachitensis, C. padifolia, C. pennsylvanica, C. persimilis, C. pexa, C. phaenopyrum, C. phippsii, C. pinetorum, C. populnea, C. prona, C. pruinosa, C. pulcherrima, C. punctata, C. purpurella, C. quaesita, C. reverchonii, C. rivularis, C. rivuloadamensis, C. rivulopugnensis, C. roribacca, C. rubella, C. rubribracteolata, C. saligna, C. sargentii, C. scabrida, C. schizophylla, C. schuettei, C. segnis, C. senta, C. sheila-phippsiae, C. sheridana, C. shuswapensis, C. sororia, C. spathulata, C. spes-aestatum, C. stolonifera, C. stonei, C. submollis, C. suborbiculata, C. succulenta, C. tecta, C. teres, C. texana, C. tracyi, C. triflora, C. turnerorum, C. uniflora, C. ursopedensis, C. venusta, C. viridis, C. visenda, C. wattiana, C. williamsii, C. wootoniana, C. ×atrorubens, C. ×bicknellii, C. ×coleae, C. ×collicola, C. ×disperma, C. ×dispessa, C. ×fretalis, C. ×incaedua, C. ×kelloggii, C. ×latebrosa, C. ×lucorum, C. ×rufula, C. ×sicca, C. ×vailiae |
C. aemula, C. aestivalis, C. alabamensis, C. alleghaniensis, C. annosa, C. aprica, C. aquacervensis, C. ashei, C. atrovirens, C. attrita, C. austromontana, C. beata, C. berberifolia, C. biltmoreana, C. brachyacantha, C. brainerdii, C. brazoria, C. brittonii, C. buckleyi, C. calpodendron, C. castlegarensis, C. chrysocarpa, C. coccinea, C. coccinioides, C. cognata, C. collina, C. colonica, C. communis, C. compacta, C. condigna, C. craytonii, C. crocea, C. crus-galli, C. cupressocollina, C. delawarensis, C. dispar, C. dodgei, C. douglasii, C. egens, C. egregia, C. enderbyensis, C. erythropoda, C. exilis, C. eximia, C. extraria, C. fecunda, C. flabellata, C. flava, C. florens, C. floridana, C. florifera, C. fluviatilis, C. formosa, C. frugiferens, C. furtiva, C. gattingeri, C. gaylussacia, C. gilva, C. greggiana, C. harbisonii, C. holmesiana, C. ignava, C. incilis, C. integra, C. intricata, C. invicta, C. iracunda, C. irrasa, C. jesupii, C. jonesiae, C. lacrimata, C. laevigata, C. lancei, C. lanuginosa, C. lassa, C. laurentiana, C. leonensis, C. lepida, C. levis, C. lumaria, C. macracantha, C. macrosperma, C. magniflora, C. margarettae, C. marshallii, C. mendosa, C. meridiana, C. mira, C. mollis, C. monogyna, C. munda, C. nananixonii, C. neobushii, C. nitida, C. oakesiana, C. okanaganensis, C. okennonii, C. opaca, C. opima, C. orbicularis, C. ouachitensis, C. padifolia, C. pennsylvanica, C. persimilis, C. pexa, C. phaenopyrum, C. phippsii, C. pinetorum, C. populnea, C. prona, C. pruinosa, C. pulcherrima, C. punctata, C. purpurella, C. quaesita, C. reverchonii, C. rivularis, C. rivuloadamensis, C. rivulopugnensis, C. roribacca, C. rubella, C. rubribracteolata, C. saligna, C. sargentii, C. scabrida, C. schizophylla, C. schuettei, C. segnis, C. senta, C. sheila-phippsiae, C. sheridana, C. shuswapensis, C. sororia, C. spathulata, C. spes-aestatum, C. stolonifera, C. stonei, C. submollis, C. suborbiculata, C. succulenta, C. tecta, C. teres, C. texana, C. tracyi, C. triflora, C. turnerorum, C. uniflora, C. ursopedensis, C. venusta, C. viridis, C. visenda, C. wattiana, C. williamsii, C. wootoniana, C. ×atrorubens, C. ×bicknellii, C. ×coleae, C. ×collicola, C. ×disperma, C. ×dispessa, C. ×fretalis, C. ×incaedua, C. ×kelloggii, C. ×latebrosa, C. ×lucorum, C. ×rufula, C. ×sicca, C. ×vailiae |
| Mespilus flabellata, C. crudelis, C. flabellata var. densiflora, C. flabellata var. grayana, C. grayana |
C. inops, C. lassa var. lanata |
| (Bosc ex Spach) K. Koch: Verh. Vereins Beförd. Gartenbaues Königl. Preuss. Staaten, ser. 2, 1: 240. (1853) |
Beadle: Biltmore Bot. Stud. 1: 86. (1902) |
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