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aubépine flabelliforme, fan-leaf hawthorn
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| Shrubs or trees, 40–50 dm. |
Shrubs or trees, 30–60 dm, branches ± weeping. |
twigs: new growth reddish at first, usually sparsely pubescent, 1-year old deep chestnut brown, older dull gray; thorns on twigs 1-year old shiny, dark brown, stout to slender, 4–6 cm. |
twigs: new growth sparsely to densely spreading-pubescent, 1-year old tawny to reddish brown, 2-years old dark gray; thorns on twigs usually numerous, straight to slightly decurved, 2-years old shiny blackish gray, ± fine to ± stout, 2–4.5 cm. |
petiole 1–2 mm wide, length 40–50% blade, glandular; blade glossy, dark green, broadly elliptic to ovate, 4–7 cm, base cuneate to slightly rounded, lobes 4 or 5 per side, sinuses deep, lobe apex acuminate, margins sharply serrate, teeth gland-tipped, veins 5 or 6 per side, apex acute, abaxial surface glabrous except along veins, adaxial densely appressed-scabrous young, glabrescent. |
petiole length 25–33% blade, ± pilose young, glandular; blade dark green mature adaxially, narrowly obovate to obovate, 2–4.5 cm, thin, slightly coriaceous, base abruptly tapered, lobes 0, or 1 or 2 per side distally, obscure, lobe apex obtuse to subacute, margins ± coarsely glandular-serrate, veins (1 or)2–4 per side, apex often ± cuspidate, adaxially shiny mature, surfaces glabrous, veins sparsely pubescent; on extension shoots broadly elliptic to ± orbiculate, larger, often slightly incised, strongly serrate. |
5–10-flowered; branches villous; bracteole frequency not recorded, linear. |
3–6-flowered; branches sparsely to moderately pilose; bracteoles caducous, linear, margins glandular. |
13–17 mm diam.; hypanthium glabrous or villous at base; sepals 4–5 mm, margins entire, glandular to glandular-serrate, rarely nearly eglandular, apex acute, abaxial pubescence not recorded; stamens 5–10 or 20, anthers usually pink; styles 3–5. |
15–18(–25) mm diam.; hypanthium sparsely pubescent; sepals narrowly triangular, 4–5 mm, margins glandular-serrate, adaxially sparsely hirsute; anthers pale yellow; styles 3(–5). |
crimson, ± ellipsoid, 8–10 mm diam., sometimes ± pruinose; sepals spreading or usually erose, not or obscurely elevated; pyrenes 3–5, dorsally deeply grooved. |
orange-red to red, suborbicular, 8–15 mm diam., glabrous; sepals spreading, 7 mm; pyrenes 3(–5). |
= 51. |
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| Flowering May–Jun; fruiting Sep–Oct. |
Flowering Mar–Apr; fruiting Aug–Sep. |
| Brush, successional fields, fencelines, woodland edges, open woodlands |
Brush on sand plains |
| 10–200 m (0–700 ft) |
0–200 m (0–700 ft) |
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CT; MA; ME; NH; NY; VT; NB; NS; ON; QC
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AL; FL; GA; LA; MS; NC; SC |
Crataegus flabellata occurs along the Niagara Escarpment in Ontario, on both sides of Lake Ontario and the St. Lawrence, to New Brunswick and Nova Scotia, and to Massachusetts. Reports for Newfoundland have not been substantiated. Crataegus flabellata is distinctive with its leaf blades glossy dark green, ovate to broadly elliptic, cuneate at the base, and with deep acuminate lobes. Specimens with less glandularity or serration of the sepal margins may resemble C. fluviatilis. In spite of the obvious differences in morphology and range, C. flabellata and C. macrosperma often have been combined by floristicians. From C. chrysocarpa, C. flabellata is distinguished by its shinier, ellipsoid, glabrous fruit, by the usually deeper, acuminate lobes of its glossy, dark green leaves, and by longer, slightly more acute sepals. Typically, the inflorescences and hypanthia of eastern C. chrysocarpa are densely hairy, practically tomentose, unlike C. flabellata with its more sparsely hairy pedicels and glabrous distal hypanthia. It is conceivable that, due to morphological intermediacy and the sympatry of all three species within the range of C. flabellata, this species represents the result of ancient hybridization between C. chrysocarpa and C. macrosperma. Crataegus flabellata is usually divided into two varieties: var. flabellata (10 stamens) and var. grayana (20 stamens), the latter being particularly common in southern Quebec. There appears to be little, if any, other constant differentiating feature. Crataegus crudelis, also from Quebec, is a form recognized by long thorns (to 10 cm). The name Crataegus densiflora Sargent, which pertains to C. flabellata, is illegitimate. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Crataegus florens occurs mainly from central Mississippi to the Carolinas and extreme northern Florida; it is common in central Alabama and is disjunct in West Feliciana Parish, Louisiana. The species may occur in more mesic habitats than other members of ser. Lacrimatae. The type form of Crataegus florens has the smallest, least strongly serrate leaves, veins 2 or 3 per side, and largest flowers, but the circumscription is here expanded to include forms of smaller flower dimensions and with larger, more strongly serrate leaves, including those with 3 or 4 veins per side. As such, C. florens is a fairly common and striking plant with its glossy, deep green leaves, thus superficially resembling members of ser. Crus-galli except for the strongly flexuous twigs. Except for its type form, which is uncommon, it flowers before most other sympatric members of ser. Lacrimatae. Differences from C. attrita, to which it is most similar, and C. teres are given in the key and discussed under those species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 9, p. 567. |
FNA vol. 9, p. 628. |
| Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Tenuifoliae |
Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Lacrimatae |
C. aemula, C. aestivalis, C. alabamensis, C. alleghaniensis, C. annosa, C. aprica, C. aquacervensis, C. ashei, C. atrovirens, C. attrita, C. austromontana, C. beata, C. berberifolia, C. biltmoreana, C. brachyacantha, C. brainerdii, C. brazoria, C. brittonii, C. buckleyi, C. calpodendron, C. castlegarensis, C. chrysocarpa, C. coccinea, C. coccinioides, C. cognata, C. collina, C. colonica, C. communis, C. compacta, C. condigna, C. craytonii, C. crocea, C. crus-galli, C. cupressocollina, C. delawarensis, C. dispar, C. dodgei, C. douglasii, C. egens, C. egregia, C. enderbyensis, C. erythropoda, C. exilis, C. eximia, C. extraria, C. fecunda, C. flava, C. florens, C. floridana, C. florifera, C. fluviatilis, C. formosa, C. frugiferens, C. furtiva, C. gattingeri, C. gaylussacia, C. gilva, C. greggiana, C. harbisonii, C. holmesiana, C. ignava, C. incilis, C. integra, C. intricata, C. invicta, C. iracunda, C. irrasa, C. jesupii, C. jonesiae, C. lacrimata, C. laevigata, C. lanata, C. lancei, C. lanuginosa, C. lassa, C. laurentiana, C. leonensis, C. lepida, C. levis, C. lumaria, C. macracantha, C. macrosperma, C. magniflora, C. margarettae, C. marshallii, C. mendosa, C. meridiana, C. mira, C. mollis, C. monogyna, C. munda, C. nananixonii, C. neobushii, C. nitida, C. oakesiana, C. okanaganensis, C. okennonii, C. opaca, C. opima, C. orbicularis, C. ouachitensis, C. padifolia, C. pennsylvanica, C. persimilis, C. pexa, C. phaenopyrum, C. phippsii, C. pinetorum, C. populnea, C. prona, C. pruinosa, C. pulcherrima, C. punctata, C. purpurella, C. quaesita, C. reverchonii, C. rivularis, C. rivuloadamensis, C. rivulopugnensis, C. roribacca, C. rubella, C. rubribracteolata, C. saligna, C. sargentii, C. scabrida, C. schizophylla, C. schuettei, C. segnis, C. senta, C. sheila-phippsiae, C. sheridana, C. shuswapensis, C. sororia, C. spathulata, C. spes-aestatum, C. stolonifera, C. stonei, C. submollis, C. suborbiculata, C. succulenta, C. tecta, C. teres, C. texana, C. tracyi, C. triflora, C. turnerorum, C. uniflora, C. ursopedensis, C. venusta, C. viridis, C. visenda, C. wattiana, C. williamsii, C. wootoniana, C. ×atrorubens, C. ×bicknellii, C. ×coleae, C. ×collicola, C. ×disperma, C. ×dispessa, C. ×fretalis, C. ×incaedua, C. ×kelloggii, C. ×latebrosa, C. ×lucorum, C. ×rufula, C. ×sicca, C. ×vailiae |
C. aemula, C. aestivalis, C. alabamensis, C. alleghaniensis, C. annosa, C. aprica, C. aquacervensis, C. ashei, C. atrovirens, C. attrita, C. austromontana, C. beata, C. berberifolia, C. biltmoreana, C. brachyacantha, C. brainerdii, C. brazoria, C. brittonii, C. buckleyi, C. calpodendron, C. castlegarensis, C. chrysocarpa, C. coccinea, C. coccinioides, C. cognata, C. collina, C. colonica, C. communis, C. compacta, C. condigna, C. craytonii, C. crocea, C. crus-galli, C. cupressocollina, C. delawarensis, C. dispar, C. dodgei, C. douglasii, C. egens, C. egregia, C. enderbyensis, C. erythropoda, C. exilis, C. eximia, C. extraria, C. fecunda, C. flabellata, C. flava, C. floridana, C. florifera, C. fluviatilis, C. formosa, C. frugiferens, C. furtiva, C. gattingeri, C. gaylussacia, C. gilva, C. greggiana, C. harbisonii, C. holmesiana, C. ignava, C. incilis, C. integra, C. intricata, C. invicta, C. iracunda, C. irrasa, C. jesupii, C. jonesiae, C. lacrimata, C. laevigata, C. lanata, C. lancei, C. lanuginosa, C. lassa, C. laurentiana, C. leonensis, C. lepida, C. levis, C. lumaria, C. macracantha, C. macrosperma, C. magniflora, C. margarettae, C. marshallii, C. mendosa, C. meridiana, C. mira, C. mollis, C. monogyna, C. munda, C. nananixonii, C. neobushii, C. nitida, C. oakesiana, C. okanaganensis, C. okennonii, C. opaca, C. opima, C. orbicularis, C. ouachitensis, C. padifolia, C. pennsylvanica, C. persimilis, C. pexa, C. phaenopyrum, C. phippsii, C. pinetorum, C. populnea, C. prona, C. pruinosa, C. pulcherrima, C. punctata, C. purpurella, C. quaesita, C. reverchonii, C. rivularis, C. rivuloadamensis, C. rivulopugnensis, C. roribacca, C. rubella, C. rubribracteolata, C. saligna, C. sargentii, C. scabrida, C. schizophylla, C. schuettei, C. segnis, C. senta, C. sheila-phippsiae, C. sheridana, C. shuswapensis, C. sororia, C. spathulata, C. spes-aestatum, C. stolonifera, C. stonei, C. submollis, C. suborbiculata, C. succulenta, C. tecta, C. teres, C. texana, C. tracyi, C. triflora, C. turnerorum, C. uniflora, C. ursopedensis, C. venusta, C. viridis, C. visenda, C. wattiana, C. williamsii, C. wootoniana, C. ×atrorubens, C. ×bicknellii, C. ×coleae, C. ×collicola, C. ×disperma, C. ×dispessa, C. ×fretalis, C. ×incaedua, C. ×kelloggii, C. ×latebrosa, C. ×lucorum, C. ×rufula, C. ×sicca, C. ×vailiae |
| Mespilus flabellata, C. crudelis, C. flabellata var. densiflora, C. flabellata var. grayana, C. grayana |
C. alabamensis var. florens |
| (Bosc ex Spach) K. Koch: Verh. Vereins Beförd. Gartenbaues Königl. Preuss. Staaten, ser. 2, 1: 240. (1853) |
Beadle: Biltmore Bot. Stud. 1: 94. (1902) |
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