Crataegus flabellata |
Crataegus eximia |
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aubépine flabelliforme, fan-leaf hawthorn |
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Habit | Shrubs or trees, 40–50 dm. | Shrubs or trees, 30–40 dm. |
Stems | twigs: new growth reddish at first, usually sparsely pubescent, 1-year old deep chestnut brown, older dull gray; thorns on twigs 1-year old shiny, dark brown, stout to slender, 4–6 cm. |
twigs: new growth reddish green, 1-year old reddish brown, older gray; thorns on twigs slightly curved, 2-years old blackish gray, usually fine, 3–4 cm. |
Leaves | petiole 1–2 mm wide, length 40–50% blade, glandular; blade glossy, dark green, broadly elliptic to ovate, 4–7 cm, base cuneate to slightly rounded, lobes 4 or 5 per side, sinuses deep, lobe apex acuminate, margins sharply serrate, teeth gland-tipped, veins 5 or 6 per side, apex acute, abaxial surface glabrous except along veins, adaxial densely appressed-scabrous young, glabrescent. |
petiole length 25–40% blade, densely stipitate-glandular (at least young); blade ovate-rhombic to ovate-trullate, 4.5–6.5 cm (flowering) to 8 cm (mature), base cuneate to broadly cuneate, lobes 3 or 4(or 5) per side, sinuses deep, lobe apex subacute to acute, angled, margins finely serrate, veins 5 or 6 per side, apex acute, adaxial midvein scabrous young. |
Inflorescences | 5–10-flowered; branches villous; bracteole frequency not recorded, linear. |
4–6-flowered; branches glabrous; bracteoles caducous, very narrowly obovate, ± membranous, margins glandular. |
Flowers | 13–17 mm diam.; hypanthium glabrous or villous at base; sepals 4–5 mm, margins entire, glandular to glandular-serrate, rarely nearly eglandular, apex acute, abaxial pubescence not recorded; stamens 5–10 or 20, anthers usually pink; styles 3–5. |
22–25 mm diam.; sepals narrowly triangular, 4–5 mm, margins finely glandular-serrate; anthers cream; styles 3 or 4. |
Pomes | crimson, ± ellipsoid, 8–10 mm diam., sometimes ± pruinose; sepals spreading or usually erose, not or obscurely elevated; pyrenes 3–5, dorsally deeply grooved. |
red, orbicular, 8–12 mm diam.; sepals spreading; pyrenes 3–5. |
2n | = 51. |
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Crataegus flabellata |
Crataegus eximia |
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Phenology | Flowering May–Jun; fruiting Sep–Oct. | Flowering late May; fruiting Sep–Oct. |
Habitat | Brush, successional fields, fencelines, woodland edges, open woodlands | Rocky woods and glades |
Elevation | 10–200 m (0–700 ft) | 200 m (700 ft) |
Distribution |
CT; MA; ME; NH; NY; VT; NB; NS; ON; QC
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AL |
Discussion | Crataegus flabellata occurs along the Niagara Escarpment in Ontario, on both sides of Lake Ontario and the St. Lawrence, to New Brunswick and Nova Scotia, and to Massachusetts. Reports for Newfoundland have not been substantiated. Crataegus flabellata is distinctive with its leaf blades glossy dark green, ovate to broadly elliptic, cuneate at the base, and with deep acuminate lobes. Specimens with less glandularity or serration of the sepal margins may resemble C. fluviatilis. In spite of the obvious differences in morphology and range, C. flabellata and C. macrosperma often have been combined by floristicians. From C. chrysocarpa, C. flabellata is distinguished by its shinier, ellipsoid, glabrous fruit, by the usually deeper, acuminate lobes of its glossy, dark green leaves, and by longer, slightly more acute sepals. Typically, the inflorescences and hypanthia of eastern C. chrysocarpa are densely hairy, practically tomentose, unlike C. flabellata with its more sparsely hairy pedicels and glabrous distal hypanthia. It is conceivable that, due to morphological intermediacy and the sympatry of all three species within the range of C. flabellata, this species represents the result of ancient hybridization between C. chrysocarpa and C. macrosperma. Crataegus flabellata is usually divided into two varieties: var. flabellata (10 stamens) and var. grayana (20 stamens), the latter being particularly common in southern Quebec. There appears to be little, if any, other constant differentiating feature. Crataegus crudelis, also from Quebec, is a form recognized by long thorns (to 10 cm). The name Crataegus densiflora Sargent, which pertains to C. flabellata, is illegitimate. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Known only from the flowering type specimen and the protologue by Beadle, Crataegus eximia seems one of the most distinct entities in the series on the basis of its large rhomboid-ovate leave with deep sinuses and angled lobes, large flowers, late flowering date (23 May in 1901). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 567. | FNA vol. 9, p. 585. |
Parent taxa | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Tenuifoliae | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Pulcherrimae |
Sibling taxa | ||
Synonyms | Mespilus flabellata, C. crudelis, C. flabellata var. densiflora, C. flabellata var. grayana, C. grayana | |
Name authority | (Bosc ex Spach) K. Koch: Verh. Vereins Beförd. Gartenbaues Königl. Preuss. Staaten, ser. 2, 1: 240. (1853) | Beadle: Biltmore Bot. Stud. 1: 62. (1902) |
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