Crataegus flabellata |
Crataegus ashei |
|
---|---|---|
aubépine flabelliforme, fan-leaf hawthorn |
Ashe hawthorn |
|
Habit | Shrubs or trees, 40–50 dm. | Shrubs, 30–50 dm. |
Stems | twigs: new growth reddish at first, usually sparsely pubescent, 1-year old deep chestnut brown, older dull gray; thorns on twigs 1-year old shiny, dark brown, stout to slender, 4–6 cm. |
twigs: new growth densely pubescent, 1-year old tan to chestnut, ± shiny, older gray; thorns on twigs few to numerous, 2-years old shiny, very dark brown to black, ± slender, 2–3 cm. |
Leaves | petiole 1–2 mm wide, length 40–50% blade, glandular; blade glossy, dark green, broadly elliptic to ovate, 4–7 cm, base cuneate to slightly rounded, lobes 4 or 5 per side, sinuses deep, lobe apex acuminate, margins sharply serrate, teeth gland-tipped, veins 5 or 6 per side, apex acute, abaxial surface glabrous except along veins, adaxial densely appressed-scabrous young, glabrescent. |
petiole length 25% blade, densely pubescent, stipitate-glandular; blade mid green, elliptic to narrowly obovate, 3–6 cm, base cuneate, lobes 0 or obscure, margins glandular-serrate, teeth to 1.5 mm, proximal teeth stipitate-glandular, veins 5–7 per side, apex acute to subacute, abaxial surface sparsely pubescent or glabrous, veins hairy, adaxial shiny, scabrous. |
Inflorescences | 5–10-flowered; branches villous; bracteole frequency not recorded, linear. |
3–10-flowered, arising proleptically (subterminal on perennial short shoots); branches densely villous; bracteoles caducous, linear, 4–8 mm, herbaceous to membranous, margins stipitate-glandular. |
Flowers | 13–17 mm diam.; hypanthium glabrous or villous at base; sepals 4–5 mm, margins entire, glandular to glandular-serrate, rarely nearly eglandular, apex acute, abaxial pubescence not recorded; stamens 5–10 or 20, anthers usually pink; styles 3–5. |
(15–)20–23 mm diam.; hypanthium tomentose; sepals narrowly triangular, 6–7 mm, margins glandular-pectinate, abaxially pilose; stamens 20(–25), anthers cream, sometimes pink; styles 3(–5) (with bristly hairs at bases). |
Pomes | crimson, ± ellipsoid, 8–10 mm diam., sometimes ± pruinose; sepals spreading or usually erose, not or obscurely elevated; pyrenes 3–5, dorsally deeply grooved. |
orange-red to deep red, 10–14 mm diam., densely short-pubescent; sepals patent-incurved; pyrenes 3(–5). |
2n | = 51. |
= 68. |
Crataegus flabellata |
Crataegus ashei |
|
Phenology | Flowering May–Jun; fruiting Sep–Oct. | Flowering Apr; fruiting Sep–Nov. |
Habitat | Brush, successional fields, fencelines, woodland edges, open woodlands | Brush, open woodlands |
Elevation | 10–200 m (0–700 ft) | 0–300 m (0–1000 ft) |
Distribution |
CT; MA; ME; NH; NY; VT; NB; NS; ON; QC
|
AL; LA; MS; TN |
Discussion | Crataegus flabellata occurs along the Niagara Escarpment in Ontario, on both sides of Lake Ontario and the St. Lawrence, to New Brunswick and Nova Scotia, and to Massachusetts. Reports for Newfoundland have not been substantiated. Crataegus flabellata is distinctive with its leaf blades glossy dark green, ovate to broadly elliptic, cuneate at the base, and with deep acuminate lobes. Specimens with less glandularity or serration of the sepal margins may resemble C. fluviatilis. In spite of the obvious differences in morphology and range, C. flabellata and C. macrosperma often have been combined by floristicians. From C. chrysocarpa, C. flabellata is distinguished by its shinier, ellipsoid, glabrous fruit, by the usually deeper, acuminate lobes of its glossy, dark green leaves, and by longer, slightly more acute sepals. Typically, the inflorescences and hypanthia of eastern C. chrysocarpa are densely hairy, practically tomentose, unlike C. flabellata with its more sparsely hairy pedicels and glabrous distal hypanthia. It is conceivable that, due to morphological intermediacy and the sympatry of all three species within the range of C. flabellata, this species represents the result of ancient hybridization between C. chrysocarpa and C. macrosperma. Crataegus flabellata is usually divided into two varieties: var. flabellata (10 stamens) and var. grayana (20 stamens), the latter being particularly common in southern Quebec. There appears to be little, if any, other constant differentiating feature. Crataegus crudelis, also from Quebec, is a form recognized by long thorns (to 10 cm). The name Crataegus densiflora Sargent, which pertains to C. flabellata, is illegitimate. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Crataegus ashei occurs from Louisiana to Alabama and is apparently scarce. Crataegus ashei has often been confused with both C. triflora (ser. Triflorae) and C. harbisonii; it differs from C. triflora in its different growth habit with branching more or less layered and leaves smallish, flowers smaller, stamens 20–24, and leaves usually more coriaceous and shiny with lobes absent. Crataegus ashei is more similar to C. harbisonii, mainly differing in being smaller in its vegetative parts, invariably lacking reproductive side shoots of the season, and by having entire leaves. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 567. | FNA vol. 9, p. 608. |
Parent taxa | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Tenuifoliae | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Bracteatae |
Sibling taxa | ||
Synonyms | Mespilus flabellata, C. crudelis, C. flabellata var. densiflora, C. flabellata var. grayana, C. grayana | |
Name authority | (Bosc ex Spach) K. Koch: Verh. Vereins Beförd. Gartenbaues Königl. Preuss. Staaten, ser. 2, 1: 240. (1853) | Beadle: Bot. Gaz. 30: 339. (1900) |
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