Crataegus flabellata |
Crataegus aemula |
|
---|---|---|
aubépine flabelliforme, fan-leaf hawthorn |
rome hawthorn |
|
Habit | Shrubs or trees, 40–50 dm. | Shrubs or trees, 30–50 dm. |
Stems | twigs: new growth reddish at first, usually sparsely pubescent, 1-year old deep chestnut brown, older dull gray; thorns on twigs 1-year old shiny, dark brown, stout to slender, 4–6 cm. |
trunks short, slender, 5–7 cm diam., bark smooth to slightly fissured or scaly; compound thorns on trunks sometimes present; twigs: new growth reddish green, pubescent, 1-year old pale tan, becoming gray-brown; thorns on twigs ± straight, chestnut brown, gray later, ± slender, 3–5 cm. |
Leaves | petiole 1–2 mm wide, length 40–50% blade, glandular; blade glossy, dark green, broadly elliptic to ovate, 4–7 cm, base cuneate to slightly rounded, lobes 4 or 5 per side, sinuses deep, lobe apex acuminate, margins sharply serrate, teeth gland-tipped, veins 5 or 6 per side, apex acute, abaxial surface glabrous except along veins, adaxial densely appressed-scabrous young, glabrescent. |
petiole 0.5–1.5 cm, pubescent at least young, stipitate-glandular; blade elliptic-ovate or broadly elliptic to suborbicular, 3–5 cm length/width = to 1.5, ± thin, base rounded or broadly truncate, lobes 3 or 4 per side, sinuses shallow, lobe apex acute, margins serrate, teeth regular, veins 4 or 5 per side, apex acute, abaxial surface glabrous, veins sparsely hairy, adaxial sparsely pubescent young, glabrescent. |
Inflorescences | 5–10-flowered; branches villous; bracteole frequency not recorded, linear. |
5–10-flowered; branches coarsely hairy; bracteole margins densely glandular. |
Flowers | 13–17 mm diam.; hypanthium glabrous or villous at base; sepals 4–5 mm, margins entire, glandular to glandular-serrate, rarely nearly eglandular, apex acute, abaxial pubescence not recorded; stamens 5–10 or 20, anthers usually pink; styles 3–5. |
14–18 mm diam.; hypanthium sparsely pubescent; sepals lanceolate, 4 mm, margins glandular-serrate, abaxial pubescence not recorded; stamens 10, anthers pink-purple; styles 3–5. |
Pomes | crimson, ± ellipsoid, 8–10 mm diam., sometimes ± pruinose; sepals spreading or usually erose, not or obscurely elevated; pyrenes 3–5, dorsally deeply grooved. |
bright red, suborbicular, 10–13 mm diam., shiny; sepals spreading-reflexed or often erose; pyrenes 3–5, dorsally shallowly grooved. |
2n | = 51. |
= 51, 68. |
Crataegus flabellata |
Crataegus aemula |
|
Phenology | Flowering May–Jun; fruiting Sep–Oct. | Flowering Apr; fruiting Sep–Oct. |
Habitat | Brush, successional fields, fencelines, woodland edges, open woodlands | Oak flatwoods, brush |
Elevation | 10–200 m (0–700 ft) | 50–600 m (200–2000 ft) |
Distribution |
CT; MA; ME; NH; NY; VT; NB; NS; ON; QC
|
AL; GA; MS |
Discussion | Crataegus flabellata occurs along the Niagara Escarpment in Ontario, on both sides of Lake Ontario and the St. Lawrence, to New Brunswick and Nova Scotia, and to Massachusetts. Reports for Newfoundland have not been substantiated. Crataegus flabellata is distinctive with its leaf blades glossy dark green, ovate to broadly elliptic, cuneate at the base, and with deep acuminate lobes. Specimens with less glandularity or serration of the sepal margins may resemble C. fluviatilis. In spite of the obvious differences in morphology and range, C. flabellata and C. macrosperma often have been combined by floristicians. From C. chrysocarpa, C. flabellata is distinguished by its shinier, ellipsoid, glabrous fruit, by the usually deeper, acuminate lobes of its glossy, dark green leaves, and by longer, slightly more acute sepals. Typically, the inflorescences and hypanthia of eastern C. chrysocarpa are densely hairy, practically tomentose, unlike C. flabellata with its more sparsely hairy pedicels and glabrous distal hypanthia. It is conceivable that, due to morphological intermediacy and the sympatry of all three species within the range of C. flabellata, this species represents the result of ancient hybridization between C. chrysocarpa and C. macrosperma. Crataegus flabellata is usually divided into two varieties: var. flabellata (10 stamens) and var. grayana (20 stamens), the latter being particularly common in southern Quebec. There appears to be little, if any, other constant differentiating feature. Crataegus crudelis, also from Quebec, is a form recognized by long thorns (to 10 cm). The name Crataegus densiflora Sargent, which pertains to C. flabellata, is illegitimate. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Crataegus aemula grows from northwestern Georgia to central Mississippi; it is locally common. The species has escaped notice in floras since Beadle in Small (1903). It is distinguished from sympatric species of the series by twig color, new growth and inflorescence indumentum, leaf shape, stipitate-glandular petioles, and bright red pomes. Remarkably, it seems most similar to C. frugiferens, provisionally placed in ser. Apricae, with which it shares stipitate-glandular petioles but differs in larger (3.5–5 cm) leaves of a proportionately narrower (somewhat narrowly ovate to broadly elliptic) shape with cuneate bases and larger marginal teeth, and 2 or 3 distinct lobes per side. Crataegus frugiferens also has a similar distribution. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 567. | FNA vol. 9, p. 569. |
Parent taxa | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Tenuifoliae | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Populneae |
Sibling taxa | ||
Synonyms | Mespilus flabellata, C. crudelis, C. flabellata var. densiflora, C. flabellata var. grayana, C. grayana | |
Name authority | (Bosc ex Spach) K. Koch: Verh. Vereins Beförd. Gartenbaues Königl. Preuss. Staaten, ser. 2, 1: 240. (1853) | Beadle: Biltmore Bot. Stud. 1: 53. (1902) |
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