Crataegus douglasii |
Crataegus aprica |
|
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black hawthorn, Douglas' hawthorn |
sunny hawthorn |
|
Habit | Shrubs, 40–80 dm. | Shrubs, 20–30 dm. |
Stems | 1-year old twigs deep tan to dark mahogany, ± shiny young, older gray to dark gray, often showing brown or alternatively tan-brown eastward; thorns on twigs straight or slightly recurved, deep brown young, (1.5–)2–3.5 cm. |
twigs ± flexuous, new growth olive green, sparsely pubescent, 1-year old reddish brown, pubescent, older dark gray-brown, glabrous; thorns on twigs straight or recurved, 2-years old dark gray-brown, slender, 3–4 cm. |
Leaves | petiole 0.7–1.5 cm, pubescent young, slightly glandular; blade usually elliptic to broadly elliptic or subrhombic (elliptic-obovate when lobes very small), 4–7 cm, lobes 2–4 per side, sinuses shallow, LII 10–20%, lobe apex subobtuse to acute, margins coarsely, sharply serrate, teeth small, gland-tipped young, venation craspedodromous, veins 4 or 5 per side, apex acute to obtuse, abaxial surface usually glabrous except on veins, adaxial densely short-appressed-pubescent. |
petiole length 20–25% blade, winged distally, pubescent, sessile-glandular; blade rhombic-elliptic or broadly elliptic, 1.5–4 cm, widest near midpoint, ± coriaceous, base sharply constricted, lobes 0 or apiculi, sinuses very shallow, lobe apex acute, margins crenate or serrate, teeth obtuse gland-tipped, veins 3 or 4 per side, apex obtuse, abaxial surface glabrous, adaxial pilose young, glabrescent. |
Inflorescences | 10–25-flowered; branches glabrous; bracteole margins stipitate- or sessile-glandular. |
3–6-flowered; branches ± densely pilose; bracteoles caducous, linear, margins glandular. |
Flowers | 10–15 mm diam.; hypanthium glabrous; sepals broadly triangular, 3–4 mm, margins sparsely glandular, adaxially glabrous; stamens 10, anthers pink; styles 3 or 4. |
13–15 mm diam.; hypanthium pilose (at least proximally); sepals narrowly triangular, 4 mm, margins glandular-serrate, midvein sometimes prominent, abaxially sparsely pubescent; stamens 10, anthers ivory to cream; styles 3–5. |
Pomes | vinous and usually waxy glaucous young, dull black mature, usually ellipsoid, 6–8 mm diam., glabrous; sepal remnants reflexed, apex obtuse; pyrenes 3 or 4, sides excavated, sometimes only shallowly. |
red or reddish orange, orbicular, 9–15 mm diam., sparsely hairy; sepals patent-reflexed; pyrenes 3–5. |
2n | = 68. |
|
Crataegus douglasii |
Crataegus aprica |
|
Phenology | Flowering May–Jun; fruiting Sep–Oct. | Flowering Apr–May; fruiting Sep–Oct. |
Habitat | Brush, old fields, fencerows, woodland edges | Open brush |
Elevation | 50–1600 m (200–5200 ft) | 20–800 m (100–2600 ft) |
Distribution |
AK; CA; ID; MI; MN; MT; OR; WA; WI; AB; BC; ON; SK
|
AL; GA; NC; SC; TN; VA |
Discussion | Crataegus douglasii occurs from southern and western British Columbia to the panhandle of Alaska and ranges to northern California and the Rocky Mountains of Idaho, Montana, and southwestern Alberta. Disjunct populations occur farther east in the Cypress Hills (Alberta and Saskatchewan), the Montana montane islands, and around the northern Great Lakes. In western North America, C. douglasii is found mainly near water in drier areas; in more mesic regions, it may occur in open woodlands. In the Great Lakes area, it mainly occurs in old pastures and on fencerows. Crataegus douglasii is distinguished from C. gaylussacia by having ten stamens and is rather variable in leaf shape and size. The species is distinguished from C. okennonii by the latter usually being taller and typically more erect as well as having a usually straight, single trunk, shorter thorns, ampulliform-orbicular fruit, and often purplish crimson fall foliage. Crataegus castlegarensis is the most similar species, and beyond possessing relatively few to many multiple thorns, it is distinguished from C. douglasii by its hairy hypanthia, pedicels, and, often, fruit, which is orbicular, plump, crimson to purple in late August, ripening to deep purple (sometimes black) at a time when sympatric C. douglasii is fully black. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Crataegus aprica is found at the southern end of the Appalachians and upper piedmont, where it may be quite common. Specimens from the Tuskegee National Forest, Macon County, Alabama, have narrower (ca. 2:1), more pointed leaves with 4 or 5 veins per side. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 512. | FNA vol. 9, p. 614. |
Parent taxa | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Douglasia > ser. Douglasianae | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Apricae |
Sibling taxa | ||
Synonyms | C. columbiana, C. tennowana | |
Name authority | Lindley: Edwards's Bot. Reg. 21: plate 1810. (1835) | Beadle: Bot. Gaz. 30: 335. (1900) |
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