Crataegus castlegarensis |
Crataegus ×collicola |
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Castlegar hawthorn, Castlegar hawthorne, hawthorn |
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Habit | Shrubs, 25–50 dm. | Shrubs or trees, 50–80 dm. |
Stems | erect; branches spreading; 1-year old twigs brown; thorns on twigs usually branched, some paired or in triads, straight to slightly recurved, dark brown with blackish tip young, 2–3 cm. |
twigs: new growth glabrous or densely pubescent young, 1-year old gray-brown, older grayish; thorns on twigs numerous, dark chestnut brown, older gray, slender, 2–8 cm. |
Leaves | petiole 0.7–1.5 cm, pubescent, eglandular; blade oblanceolate to ovate-rhombic, 3.5–6 cm, lobes 3 or 4 per side, sinuses shallow to deep, lobe apex usually acute, margins serrate, teeth apices finally glandular young, venation craspedodromous, veins 4 or 5 per side, apex broadly triangular, abaxial surface sparsely hairy or glabrous except on veins, adaxial conspicuously appressed-pubescent young, glabrescent except on midvein. |
petiole length 20–25% blade, pubescent, glandular; blade narrowly obovate, 3.5–5 cm, base narrowly cuneate, lobes 0, or 2 or 3 per side, sinuses shallow, lobe apex acute, margins serrulate, teeth 0.5–0.7 mm, venation camptodromous, veins 3–6 per side, apex acute, ± lustrous, abaxial surface villous on veins, adaxial soft-white-hairy young, becoming sparsely scabrous. |
Inflorescences | 12–20-flowered; branches sparsely to densely pubescent; bracteole margins stipitate-glandular. |
6–14-flowered; branches glabrous or sparsely pubescent; bracteoles caducous, numerous, reddish, very narrow, membranous, margins glandular. |
Flowers | 12 mm diam.; hypanthium pubescent or glabrous; sepals triangular, 3 mm, margins remotely glandular-serrate; stamens 10, anthers pink; styles 3 or 4. |
18 mm diam.; hypanthium densely villous; sepals narrowly triangular, margins entire or sparsely glandular-serrate; stamens 20, anthers rose; styles 3–5. |
Pomes | crimson (mid Aug) turning to reddish plum or, ultimately, blackish purple, orbicular, ± oblate (recessed at junction with pedicel), 10 mm diam., sparsely pilose; sepals reflexed, apex obtuse; pyrenes 3 or 4, sides usually pitted. |
orange to dull-red or dark red, depressed orbicular to orbicular, 8–13 mm diam., pubescent young; sepals small, erose or ± patent; pyrenes 3–5, dorsally grooved, sides plane. |
2n | = 68. |
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Crataegus castlegarensis |
Crataegus ×collicola |
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Phenology | Flowering May–Jun; fruiting Sep–Oct. | Flowering May; fruiting Sep–Oct. |
Habitat | Mesic brush | Fertile ground along streams, fields, dry pastures in mountains |
Elevation | 300–1200 m (1000–3900 ft) | 100–1000 m (300–3300 ft) |
Distribution |
CA; ID; MT; OR; UT; WA; WY; AB; BC; SK
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AR; KY; MO; NC; VA |
Discussion | Crataegus castlegarensis occurs from around the northern Okanagan, British Columbia, to the northern California Coast Ranges, to northwestern Montana, and the Rocky Mountains to northeast of Salt Lake City, Utah. The species occurs also in the Cypress Hills of Alberta and Saskatchewan; it is found in habitats similar to those of C. douglasii and is at least as abundant as that species in a number of parts of its range. Crataegus castlegarensis is readily recognized by a combination of hairy inflorescence branches, pomes more or less orbicular, crimson or burgundy (with irregular earlier ripening) around the third week of August, soon becoming purple, often when nearby C. douglasii is already black, as well as a tendency to possess thorns on the young twigs branched at the base to become double, triple, or even sometimes quadruple. Such multiple thorns, though sometimes abundant on a bush, are more often few and may require searching for. Inflorescence pubescence, as in other species with this characteristic, may become sparse by fruiting. Crataegus castlegarensis is most similar to C. douglasii; its fruit is usually more orbicular, even oblately so, than is normal in that species, sometimes even with a recessed junction to the pedicel like an apple. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Crataegus ×collicola and C. ×verruculosa are two different putative hybrids, both of which are expected to possess parentage suggested by E. J. Palmer (1952) of C. collina and C. crus-galli. Crataegus ×collicola has thorns 4–8 mm, young twigs glabrous, leaf blade broadly elliptic to obovate, 2 or 3 lobes, and 4–6 veins per side, glabrous inflorescence branches, entire sepal margins, and pomes orange to dull red, depressed orbicular, and 8–11 mm diam. Crataegus ×verruculosa has thorns 2–2.5 mm, young twigs hairy, leaf blades narrowly obovate, unlobed, veins 3–5 per side, hairy inflorescence branches, sparingly glandular-serrate sepal margins, and pomes dark red, orbicular, 10–13 mm diam. If these are considered to be the same hybrid species, separate nothovariety names should be erected to maintain distinction. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 513. | FNA vol. 9, p. 637. |
Parent taxa | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Douglasia > ser. Douglasianae | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus |
Sibling taxa | ||
Synonyms | C. collina var. collicola, C. ×verruculosa | |
Name authority | J. B. Phipps & O'Kennon: Sida 20: 121, figs. 3, 4. (2002) | Crataegus ×collicola Ashe: J. Elisha Mitchell Sci. Soc. 16: 75. (1900) — as species |
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