Crataegus castlegarensis |
Crataegus cognata |
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Castlegar hawthorn, Castlegar hawthorne, hawthorn |
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Habit | Shrubs, 25–50 dm. | Shrubs, 20–40 dm. |
Stems | erect; branches spreading; 1-year old twigs brown; thorns on twigs usually branched, some paired or in triads, straight to slightly recurved, dark brown with blackish tip young, 2–3 cm. |
main branches often suberect; twigs: new growth reddish brown, 1-year old purple-brown, older dark gray; thorns on twigs numerous, straight to slightly recurved, 2-years old black, shiny, ± slender, 3–5 cm. |
Leaves | petiole 0.7–1.5 cm, pubescent, eglandular; blade oblanceolate to ovate-rhombic, 3.5–6 cm, lobes 3 or 4 per side, sinuses shallow to deep, lobe apex usually acute, margins serrate, teeth apices finally glandular young, venation craspedodromous, veins 4 or 5 per side, apex broadly triangular, abaxial surface sparsely hairy or glabrous except on veins, adaxial conspicuously appressed-pubescent young, glabrescent except on midvein. |
petiole length 28–36% blade, eglandular or with 1–2 glands distally; blade ovate, 3.5–5.5 cm, subcoriaceous, base broadly cuneate to truncate, lobes 3(or 4) per side, sinuses shallow, lobe apex subacute, margins crenate-serrate, veins 5–7 per side, apex subacute, surfaces glabrous. |
Inflorescences | 12–20-flowered; branches sparsely to densely pubescent; bracteole margins stipitate-glandular. |
4- or 5-flowered; branches glabrous; bracteole margins short-stipitate-glandular. |
Flowers | 12 mm diam.; hypanthium pubescent or glabrous; sepals triangular, 3 mm, margins remotely glandular-serrate; stamens 10, anthers pink; styles 3 or 4. |
17–19 mm diam.; hypanthium glabrous; sepals narrowly triangular, 5 mm, base broad, margins subentire, abaxially glabrous; stamens 20, anthers white, 0.6–0.8 mm; styles 3 or 4. |
Pomes | crimson (mid Aug) turning to reddish plum or, ultimately, blackish purple, orbicular, ± oblate (recessed at junction with pedicel), 10 mm diam., sparsely pilose; sepals reflexed, apex obtuse; pyrenes 3 or 4, sides usually pitted. |
green with pink or purple pruinosity, becoming orange-red or crimson, 8–10 mm diam.; flesh thin; sepals on collar, spreading-reflexed; pyrenes 3 or 4. |
2n | = 68. |
= 51. |
Crataegus castlegarensis |
Crataegus cognata |
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Phenology | Flowering May–Jun; fruiting Sep–Oct. | Flowering Apr–May; fruiting Sep–Oct. |
Habitat | Mesic brush | Successional fields, fencerows, brush, open wood margins |
Elevation | 300–1200 m (1000–3900 ft) | 50–300 m (200–1000 ft) |
Distribution |
CA; ID; MT; OR; UT; WA; WY; AB; BC; SK
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MA; MI; MO; NY; ON |
Discussion | Crataegus castlegarensis occurs from around the northern Okanagan, British Columbia, to the northern California Coast Ranges, to northwestern Montana, and the Rocky Mountains to northeast of Salt Lake City, Utah. The species occurs also in the Cypress Hills of Alberta and Saskatchewan; it is found in habitats similar to those of C. douglasii and is at least as abundant as that species in a number of parts of its range. Crataegus castlegarensis is readily recognized by a combination of hairy inflorescence branches, pomes more or less orbicular, crimson or burgundy (with irregular earlier ripening) around the third week of August, soon becoming purple, often when nearby C. douglasii is already black, as well as a tendency to possess thorns on the young twigs branched at the base to become double, triple, or even sometimes quadruple. Such multiple thorns, though sometimes abundant on a bush, are more often few and may require searching for. Inflorescence pubescence, as in other species with this characteristic, may become sparse by fruiting. Crataegus castlegarensis is most similar to C. douglasii; its fruit is usually more orbicular, even oblately so, than is normal in that species, sometimes even with a recessed junction to the pedicel like an apple. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Crataegus cognata in the strict sense occurs from Michigan to Massachusetts with the southern limits poorly understood. Specimens from Missouri with large (5–7 cm) leaf blades and no or only weakly developed lobes should be checked against C. pruinosa var. magnifolia. Crataegus perjucunda Sargent, native to Middlesex County, Ontario, is a distinctive local species similar to C. cognata but with shorter (2–3 cm) and finer thorns as well as relatively wider leaves (length/width = 1.2). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 513. | FNA vol. 9, p. 577. |
Parent taxa | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Douglasia > ser. Douglasianae | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Pruinosae |
Sibling taxa | ||
Synonyms | C. aperta, C. patrum, C. pruinosa var. cognata, C. pruinosa var. latisepala | |
Name authority | J. B. Phipps & O'Kennon: Sida 20: 121, figs. 3, 4. (2002) | Sargent: Rhodora 5: 58. (1903) |
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