Crataegus berberifolia |
Crataegus ×disperma |
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barberry hawthorn, barberry-leaf hawthorn |
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Habit | Shrubs or trees, 60 dm. | Shrubs or trees, 50–80 dm. | ||||
Stems | twigs: new growth orange-brown or green tinged with red, ± pubescent, 1-year old brown, older gray; thorns on twigs ± straight or recurved, 2-years old shiny black to chestnut brown, fine or stouter, (2–)3–4(–6.5) cm. |
twigs: new growth reddish young, glabrous or pubescent, 1-year old shiny brown, glabrous, older dark gray; thorns on twigs 2-years old shiny, dark brown, fine, 3–4 cm. |
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Leaves | petiole 4–6 mm, length 13–18% blade, glabrescent, eglandular; blade narrowly obovate to oblanceolate, (2.5–)3(–4) cm, coriaceous, base narrowly cuneate, lobes 0, margins finely crenate or serrate except at base, or only beyond widest part, venation craspedodromous, veins 4–6 per side, apex subacute to obtuse, lustrous, abaxial surface ± densely pilose on veins, sometimes pubescent on surface, adaxial hairy young, glabrescent or becoming scabrous. |
deciduous; petiole 0.7–1.5 cm, winged nearly to base, glabrous; blade dark green, obovate or elliptic, 2.5–5 cm, subcoriaceous, base broadly to narrowly cuneate, lobes 0 or few, sinuses very shallow, margins serrate, venation craspedodromous, veins 5–7 per side, apex acute, surfaces glabrous. |
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Inflorescences | 8–12-flowered; branches densely pubescent; bracteoles linear, margins glandular. |
5–12-flowered; branches glabrous; bracteoles caducous, linear, membranous, margins glandular. |
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Flowers | 10–20 mm diam.; hypanthium villous or glabrous; sepals 3–5 mm, margins entire, abaxially glabrous; stamens 10 or 20, anthers cream or pink; styles 2 or 3. |
13–18 mm diam.; hypanthium glabrous; sepals narrow, margins ± entire or subserrate; stamens 10, anthers pink or red-purple; styles 2(or 3). |
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Pomes | reddish to yellow, suborbicular, 8–10 mm diam., glabrous; sepals erose or patent; pyrenes 2 or 3. |
deep red, ± oblong, 12–14 × 9–11 mm; sepals spreading, subsessile; pyrenes 2, dorsally grooved, sides plane. |
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Crataegus berberifolia |
Crataegus ×disperma |
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Phenology | Flowering May; fruiting Sep–Oct. | |||||
Habitat | Brush, along streams | |||||
Elevation | 50–200 m (200–700 ft) | |||||
Distribution |
AL; AR; FL; GA; KS; LA; MO; MS; NC; SC; TN
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IA; IL; IN; OH; PA; VA |
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Discussion | Varieties 2 (2 in the flora). Crataegus berberifolia is widespread from Texas to Missouri, Florida, and Virginia; it is particularly abundant in Louisiana. Crataegus berberifolia is little differentiated from some forms of C. crus-galli, except in indumentum; it has relatively small and less variably shaped leaves. Its distribution is quite different. Intermediates with hairy leaves and glabrous inflorescences or nearly glabrous leaves and hairy inflorescences may represent hybrids with C. crus-galli (where they might be reached in the key). Such a situation is found in C. araioclada. Abrasion of the adaxial leaf pubescence may occur, rendering identification more difficult with fruiting material. Crataegus berberifolia has a plethora of yellow and orange-fruited forms, particularly from southern Louisiana. Crataegus fera and C. tersa are red-fruited, C. crocina yellow. The fruit color in the type is unknown; E. J. Palmer (in specimen annotation) called it yellow-orange. A form with exceptionally white-tomentose leaves near Copenhagen, Louisiana, is probably this species. Crataegus regalis var. paradoxa (Sargent) E. J. Palmer, from Missouri and adjacent Kansas and Arkansas, is probably a hybrid between the deeply serrated 'regalis' leaf form of C. crus-galli and a form of C. berberifolia. Two common forms of C. berberifolia occur, treated here as varieties: var. engelmannii with ten pink anthers and var. berberifolia with 20 cream anthers. Forms with 20 pink or ten cream anthers also occur sporadically. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Crataegus ×disperma has been found sporadically from Iowa east to Virginia and Pennsylvania, where the ranges of C. punctata and C. crus-galli overlap. Crataegus ×disperma belongs to a group of taxa intermediate between C. crus-galli and C. punctata; it may key to C. collina, although it has glossier leaves and usually fewer styles and pyrenes. Some specimens are more hairy, or with more strongly glandular-serrate sepal margins, than in the description, which is based on Ashe. Crataegus danielsii E. J. Palmer from Missouri, with glossy, more or less coriaceous, but lobed leaves is another problematic C. crus-galli × C. collina hybrid according to Palmer (1952). The pink-anthered C. ×disperma and cream-anthered C. ×incaedua are similar to C. collina var. sordida except for the number of styles and pyrenes (two or three). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 540. | FNA vol. 9, p. 637. | ||||
Parent taxa | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Crus-galli | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | C. grandis, C. pausiaca, C. peoriensis, C. punctata var. pausiaca | |||||
Name authority | Torrey & A. Gray: Fl. N. Amer. 1: 469. (1840) | Crataegus ×disperma Ashe: J. Elisha Mitchell Sci. Soc. 17: 14. (1900) — as species | ||||
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