Crataegus berberifolia |
Crataegus schuettei |
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barberry hawthorn, barberry-leaf hawthorn |
aubépine de Schuette, royal hawthorn, Schuette hawthorn, Schuette's hawthorn |
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Habit | Shrubs or trees, 60 dm. | Shrubs or trees, 40–70 dm. | ||||||||||||||||
Stems | twigs: new growth orange-brown or green tinged with red, ± pubescent, 1-year old brown, older gray; thorns on twigs ± straight or recurved, 2-years old shiny black to chestnut brown, fine or stouter, (2–)3–4(–6.5) cm. |
twigs: new growth reddish, glabrous, 1-year old shiny reddish brown, older dull gray; thorns on twigs 1-year old glossy, dark brown to blackish, ± stout, 3–6 cm. |
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Leaves | petiole 4–6 mm, length 13–18% blade, glabrescent, eglandular; blade narrowly obovate to oblanceolate, (2.5–)3(–4) cm, coriaceous, base narrowly cuneate, lobes 0, margins finely crenate or serrate except at base, or only beyond widest part, venation craspedodromous, veins 4–6 per side, apex subacute to obtuse, lustrous, abaxial surface ± densely pilose on veins, sometimes pubescent on surface, adaxial hairy young, glabrescent or becoming scabrous. |
petiole length 50–60% blade, usually glandular; blade ovate to oblong-ovate or narrowly ovate (ovate on extension shoots), 3–6 cm, base truncate, rounded, or cuneate, lobes 4 or 5 per side, sinuses moderately deep, lobe apex acute to acuminate, margins serrate, teeth 1 mm, veins 5 or 6 per side, apex acute, adaxial surface sparsely to densely scabrous-pubescent young, early glabrescent. |
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Inflorescences | 8–12-flowered; branches densely pubescent; bracteoles linear, margins glandular. |
4–10-flowered; branches usually glabrous, rarely sparsely pubescent; bracteoles absent or few, linear. |
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Flowers | 10–20 mm diam.; hypanthium villous or glabrous; sepals 3–5 mm, margins entire, abaxially glabrous; stamens 10 or 20, anthers cream or pink; styles 2 or 3. |
15–26 mm diam.; hypanthium glabrous; sepals 3–4 mm, margins subentire or glandular-serrate, abaxially glabrous; stamens 20, anthers red; styles 3–5. |
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Pomes | reddish to yellow, suborbicular, 8–10 mm diam., glabrous; sepals erose or patent; pyrenes 2 or 3. |
red, ellipsoid to suborbicular, 8–20 mm diam., succulent; sepals ± erose or erect-patent; pyrenes 3–5, dorsally grooved. |
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2n | = 51. |
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Crataegus berberifolia |
Crataegus schuettei |
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Distribution |
AL; AR; FL; GA; KS; LA; MO; MS; NC; SC; TN
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AR; CT; IL; IN; KY; MA; MD; ME; MI; MN; MO; NC; NH; NJ; NY; OH; PA; RI; TN; VA; VT; WI; WV; ON; QC
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Discussion | Varieties 2 (2 in the flora). Crataegus berberifolia is widespread from Texas to Missouri, Florida, and Virginia; it is particularly abundant in Louisiana. Crataegus berberifolia is little differentiated from some forms of C. crus-galli, except in indumentum; it has relatively small and less variably shaped leaves. Its distribution is quite different. Intermediates with hairy leaves and glabrous inflorescences or nearly glabrous leaves and hairy inflorescences may represent hybrids with C. crus-galli (where they might be reached in the key). Such a situation is found in C. araioclada. Abrasion of the adaxial leaf pubescence may occur, rendering identification more difficult with fruiting material. Crataegus berberifolia has a plethora of yellow and orange-fruited forms, particularly from southern Louisiana. Crataegus fera and C. tersa are red-fruited, C. crocina yellow. The fruit color in the type is unknown; E. J. Palmer (in specimen annotation) called it yellow-orange. A form with exceptionally white-tomentose leaves near Copenhagen, Louisiana, is probably this species. Crataegus regalis var. paradoxa (Sargent) E. J. Palmer, from Missouri and adjacent Kansas and Arkansas, is probably a hybrid between the deeply serrated 'regalis' leaf form of C. crus-galli and a form of C. berberifolia. Two common forms of C. berberifolia occur, treated here as varieties: var. engelmannii with ten pink anthers and var. berberifolia with 20 cream anthers. Forms with 20 pink or ten cream anthers also occur sporadically. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 4 (4 in the flora). Crataegus schuettei ranges from Wisconsin through the southern Great Lakes to southern New England, to North Carolina. Recent discoveries add the Ozarks and may add Minnesota (see discussion under C. fluviatilis) to the documented range. Variety schuettei closely resembles C. macrosperma, differing principally in its often larger flowers, 20 stamens, usually glandular-serrate sepal margins, and larger fruit but is not nearly so variable as C. macrosperma; the other varieties are very rare. Still, the four varieties are fairly distinct. Crataegus basilica appears to be the southern form of the type variety and is very like sympatric C. macrosperma except for stamen number. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 540. | FNA vol. 9, p. 565. | ||||||||||||||||
Parent taxa | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Crus-galli | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Tenuifoliae | ||||||||||||||||
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Name authority | Torrey & A. Gray: Fl. N. Amer. 1: 469. (1840) | Ashe: J. Elisha Mitchell Sci. Soc. 17: 7. (1901) | ||||||||||||||||
Web links |